World's Best Scientists 2026 revealed!

D-Index & Metrics

Biology and Biochemistry

D-Index
122
Citations
56092
World Ranking
582
National Ranking
372

Medicine

D-Index
122
Citations
56023
World Ranking
3472
National Ranking
1908

Timothy Hla publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Timothy Hla sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 337 publications — 84th percentile

84% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Timothy Hla D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Timothy Hla sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 122 D-Index — 97th percentile

97% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Research.com Recognitions

  • 2016 - Fellow of the American Association for the Advancement of Science (AAAS)

Overview

Timothy Hla is affiliated with Harvard University in the United States and has contributed extensively to the fields of Biochemistry, Genetics, and Molecular Biology, with additional work in Medicine. Their research focuses primarily on Molecular Biology, Cell Biology, Immunology, Genetics, and Physiology.

The core scientific work of Timothy Hla includes studies in sphingolipid metabolism and signaling, notably within the context of vascular biology and immunological regulation. Other notable research topics involve caveolin-1 and cellular processes, zebrafish biomedical research applications, retinal diseases and treatments, phagocytosis and immune regulation, lipid metabolism and disorders, and connective tissue disorders.

Frequent publication venues for Timothy Hla include:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Journal of Biological Chemistry
  • Arteriosclerosis Thrombosis and Vascular Biology
  • JCI Insight
  • Developmental Cell

Their publication record highlights several recent papers, such as:

  • Endothelial S1P 1 Signaling Counteracts Infarct Expansion in Ischemic Stroke, 2020, Circulation Research
  • Lysophospholipid Mediators in Health and Disease, 2021, Annual Review of Pathology Mechanisms of Disease
  • Endothelial sphingosine 1-phosphate receptors promote vascular normalization and antitumor therapy, 2020, Proceedings of the National Academy of Sciences
  • S1PR1 regulates the quiescence of lymphatic vessels by inhibiting laminar shear stress-dependent VEGF-C signaling, 2020, JCI Insight
  • Sphingosine 1-phosphate receptor-targeted therapeutics in rheumatic diseases, 2022, Nature Reviews Rheumatology

Timothy Hla has collaborated frequently with colleagues including Andrew Kuo, Andréane Cartier, Colin Niaudet, B. Jung, and Steven Swendeman.

In recognition of their scientific contributions, Timothy Hla was named a Fellow of the American Association for the Advancement of Science (AAAS) in 2016.

Best Publications

  • Cyclooxygenase Isozymes: The Biology of Prostaglandin Synthesis and Inhibition

    Daniel L. Simmons;Regina M. Botting;Timothy Hla

  • Human cyclooxygenase-2 cDNA.

    Timothy Hla;Karen Neilson

  • Expression of Cyclooxygenase-1 and -2 in Human Colorectal Cancer

    Hajime Sano;Yutaka Kawahito;Ronald L. Wilder;Akira Hashiramoto

  • Edg-1, the G protein–coupled receptor for sphingosine-1-phosphate, is essential for vascular maturation

    Yujing Liu;Ryuichi Wada;Tadashi Yamashita;Yide Mi

  • Sphingosine-1-Phosphate as a Ligand for the G Protein-Coupled Receptor EDG-1

    Menq Jer Lee;James R. Van Brocklyn;James R. Van Brocklyn;James R. Van Brocklyn;Shobha Thangada;Shobha Thangada;Shobha Thangada;Catherine H. Liu;Catherine H. Liu;Catherine H. Liu

  • Vascular Endothelial Cell Adherens Junction Assembly and Morphogenesis Induced by Sphingosine-1-Phosphate

    Menq-Jer Lee;Shobha Thangada;Kevin P Claffey;Nicolas Ancellin

  • Overexpression of Cyclooxygenase-2 Is Sufficient to Induce Tumorigenesis in Transgenic Mice

    Catherine H. Liu;Sung-Hee Chang;Kirsi Narko;Ovidiu C. Trifan

  • Cyclooxygenase-1 and -2 expression in rheumatoid synovial tissues. Effects of interleukin-1 beta, phorbol ester, and corticosteroids.

    Leslie J. Crofford;Ronald L. Wilder;Ari P. Ristimaki;Hajime Sano

  • Interleukin 1 regulates synthesis of amyloid beta-protein precursor mRNA in human endothelial cells.

    D Goldgaber;H W Harris;T Hla;T Maciag

  • Structure of the human cyclo-oxygenase-2 gene

    S B Appleby;A Ristimäki;K Neilson;K Narko

  • Induction of vascular endothelial growth factor expression in synovial fibroblasts by prostaglandin E and interleukin‐1: a potential mechanism for inflammatory angiogenesis

    Pazit Ben-Av;Leslie J. Crofford;Ronald L. Wilder;Timothy Hla

  • The nuclear receptor PPAR gamma and immunoregulation: PPAR gamma mediates inhibition of helper T cell responses.

    Robert B. Clark;David Bishop-Bailey;Tatiana Estrada-Hernandez;Timothy Hla

  • Lysophospholipids--receptor revelations.

    Timothy Hla;Menq-Jer Lee;Nicolas Ancellin;Ji H. Paik

  • FTY720: sphingosine 1-phosphate receptor-1 in the control of lymphocyte egress and endothelial barrier function.

    Volker Brinkmann;Jason G. Cyster;Timothy Hla

  • Endothelial cell apoptosis induced by the peroxisome proliferator-activated receptor (PPAR) ligand 15-deoxy-Delta12, 14-prostaglandin J2.

    David Bishop-Bailey;Timothy Hla

  • Endothelium-protective sphingosine-1-phosphate provided by HDL-associated apolipoprotein M.

    Christina Christoffersen;Hideru Obinata;Sunil B. Kumaraswamy;Sylvain Galvani

  • Differential inhibition of human prostaglandin endoperoxide H synthases-1 and -2 by nonsteroidal anti-inflammatory drugs.

    O Laneuville;D K Breuer;D L Dewitt;T Hla

  • Induction of cyclooxygenase-2 by interleukin-1 alpha. Evidence for post-transcriptional regulation.

    A Ristimäki;S Garfinkel;J Wessendorf;T Maciag

  • International Union of Pharmacology. XXXIV. Lysophospholipid Receptor Nomenclature

    Jerold Chun;Edward J. Goetzl;Timothy Hla;Yasuyuki Igarashi

  • An abundant transcript induced in differentiating human endothelial cells encodes a polypeptide with structural similarities to G-protein-coupled receptors.

    T Hla;T Maciag

Frequent Co-Authors

Richard L. Proia
Richard L. Proia National Institutes of Health
Jihye Paik
Jihye Paik Cornell University
Thomas Maciag
Thomas Maciag Maine Medical Center
Kevin P. Claffey
Kevin P. Claffey University of Connecticut Health Center
Ari Ristimäki
Ari Ristimäki University of Helsinki
Andrew J. Dannenberg
Andrew J. Dannenberg Cornell University
Todd Evans
Todd Evans Cornell University
Christer Betsholtz
Christer Betsholtz Uppsala University
Kotha Subbaramaiah
Kotha Subbaramaiah Cornell University
Frederick R. Maxfield
Frederick R. Maxfield Cornell University

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