World's Best Scientists 2026 revealed!
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Rama Khokha

D-Index & Metrics

Discipline name D-Index World Ranking Current World Ranking National Ranking Current National Ranking Publications Citations
Medicine 92 11414 10730 458 433 210 27438
Biology and Biochemistry 92 2236 2033 62 57 199 27267

Rama Khokha publications per year

The chart shows the history of publications by Rama Khokha between 1984 and 2026, highlighting the no. of papers published in each year and offering an overview of the publication velocity of this scholar. Rama Khokha published across 43 years, from 1984 to 2026, averaging 7 papers a year. Output peaked at 68 publications in 2023. 14 of the 302 publications appeared in the last two years.

No. of publications
20 40 60
Bar chart. Horizontal axis: year, 1984 to 2026. Vertical axis: number of publications, 0 to 68. Peak 68 publications in 2023. 1984: 1 publication 1985: 0 publications 1986: 0 publications 1987: 2 publications 1988: 1 publication 1989: 3 publications 1990: 1 publication 1991: 2 publications 1992: 2 publications 1993: 1 publication 1994: 3 publications 1995: 4 publications 1996: 3 publications 1997: 3 publications 1998: 4 publications 1999: 4 publications 2000: 10 publications 2001: 7 publications 2002: 1 publication 2003: 11 publications 2004: 10 publications 2005: 11 publications 2006: 7 publications 2007: 9 publications 2008: 10 publications 2009: 3 publications 2010: 10 publications 2011: 5 publications 2012: 8 publications 2013: 10 publications 2014: 6 publications 2015: 12 publications 2016: 2 publications 2017: 5 publications 2018: 5 publications 2019: 12 publications 2020: 8 publications 2021: 8 publications 2022: 9 publications 2023: 68 publications 2024: 7 publications 2025: 12 publications 2026: 2 publications
1984 2026

302 publications in total across all disciplines

View publications per year as a table
Rama Khokha: publications per year, 1984 to 2026
Year Publications
1984 1
1985 0
1986 0
1987 2
1988 1
1989 3
1990 1
1991 2
1992 2
1993 1
1994 3
1995 4
1996 3
1997 3
1998 4
1999 4
2000 10
2001 7
2002 1
2003 11
2004 10
2005 11
2006 7
2007 9
2008 10
2009 3
2010 10
2011 5
2012 8
2013 10
2014 6
2015 12
2016 2
2017 5
2018 5
2019 12
2020 8
2021 8
2022 9
2023 68
2024 7
2025 12
2026 2
Total 302
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Rama Khokha publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Rama Khokha sits on this spectrum.

No. of scientists
200 400 600 800
Bar chart with 100 bars. Horizontal axis: publications, 47–56 to 1,028+. Vertical axis: number of scientists, 0 to 980. Most scientists, 980, have 127–136 publications. The last bar groups every scientist with 1,028 publications or more. The highlighted bar, 197–206 publications, is where this scientist sits. 47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 413 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 848 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 949 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 840 scientists 207–216 publications: 733 scientists 217–226 publications: 708 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 417 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 196 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 59 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientist 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientist 1,028+ publications: 100 scientists
47–56 publications 1,028+

This scientist: 199 publications — 49th percentile

49% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

View publications distribution as a table
Number of Biology and Biochemistry scientists by publication count, Research.com 2026 ranking edition. Based on 19,587 ranked scientists.
Publications Scientists This scientist
47–56 8
57–66 35
67–76 106
77–86 231
87–96 413
97–106 546
107–116 704
117–126 848
127–136 980
137–146 942
147–156 969
157–166 949
167–176 951
177–186 915
187–196 787
197–206 840 199
207–216 733
217–226 708
227–236 651
237–246 605
247–256 510
257–266 524
267–276 434
277–286 417
287–296 350
297–306 363
307–316 315
317–326 296
327–336 261
337–346 240
347–356 219
357–366 196
367–376 154
377–386 161
387–396 155
397–406 145
407–416 124
417–426 112
427–436 132
437–446 116
447–456 99
457–466 81
467–476 91
477–486 80
487–496 80
497–506 59
507–516 36
517–526 46
527–536 54
537–546 44
547–556 43
557–566 43
567–576 42
577–586 25
587–596 34
597–606 23
607–616 33
617–626 31
627–636 27
637–646 25
647–656 28
657–666 34
667–676 18
677–686 16
687–696 10
697–706 12
707–716 21
717–726 12
727–736 12
737–746 10
747–756 7
757–766 13
767–776 15
777–786 13
787–796 9
797–806 9
807–816 7
817–826 4
827–836 9
837–846 7
847–856 3
857–866 5
867–876 5
877–886 11
887–896 3
897–906 4
907–916 7
917–926 5
927–936 6
937–946 6
947–956 3
957–966 7
967–976 2
977–986 2
987–996 1
997–1,006 5
1,007–1,016 2
1,017–1,026 2
1,027 1
1,028+ 100
Download as CSV

Rama Khokha D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Rama Khokha sits on this spectrum.

No. of scientists
250 500 750 1,000 1,250
Bar chart with 65 bars. Horizontal axis: D-Index, 40–41 to 167+. Vertical axis: number of scientists, 0 to 1,253. Most scientists, 1,253, have 58–59 D-Index. The last bar groups every scientist with 167 D-Index or more. The highlighted bar, 92–93 D-Index, is where this scientist sits. 40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 316 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 899 scientists 52–53 D-Index: 1,025 scientists 54–55 D-Index: 1,149 scientists 56–57 D-Index: 1,235 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,162 scientists 62–63 D-Index: 1,130 scientists 64–65 D-Index: 1,031 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 714 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 71 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40–41 D-Index 167+

This scientist: 92 D-Index — 89th percentile

89% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

View D-Index distribution as a table
Number of Biology and Biochemistry scientists by D-index, Research.com 2026 ranking edition. Based on 19,587 ranked scientists.
D-Index Scientists This scientist
40–41 80
42–43 183
44–45 316
46–47 504
48–49 718
50–51 899
52–53 1,025
54–55 1,149
56–57 1,235
58–59 1,253
60–61 1,162
62–63 1,130
64–65 1,031
66–67 897
68–69 814
70–71 714
72–73 709
74–75 596
76–77 512
78–79 473
80–81 412
82–83 373
84–85 358
86–87 285
88–89 273
90–91 227
92–93 208 92
94–95 193
96–97 153
98–99 157
100–101 148
102–103 120
104–105 113
106–107 100
108–109 86
110–111 67
112–113 71
114–115 73
116–117 64
118–119 53
120–121 60
122–123 54
124–125 43
126–127 38
128–129 49
130–131 26
132–133 18
134–135 23
136–137 32
138–139 32
140–141 27
142–143 19
144–145 22
146–147 12
148–149 16
150–151 14
152–153 10
154–155 13
156–157 10
158–159 7
160–161 9
162–163 13
164–165 4
166 4
167+ 98
Download as CSV

Research.com Recognitions

  • 2020 - Fellow of the Royal Society of Canada Academy of Science
  • 2014 - Robert L. Noble Prize, Canadian Cancer Society

Overview

Rama Khokha is affiliated with the Princess Margaret Cancer Centre in Canada and has contributed extensively to research in oncology, molecular biology, and medicine. Their work focuses on cancer biology, particularly pancreatic and hepatic oncology, cancer genomics, epigenetics, and immunology.

The scientist's recent publications illustrate key themes across tumor microenvironments, immune regulation, and cancer subtype differentiation. Notable papers include:

  • Tryptophan-derived microbial metabolites activate the aryl hydrocarbon receptor in tumor-associated macrophages to suppress anti-tumor immunity, 2022, published in Immunity
  • Spatially confined sub-tumor microenvironments in pancreatic cancer, 2021, published in Cell
  • GATA6 Expression Distinguishes Classical and Basal-like Subtypes in Advanced Pancreatic Cancer, 2020, published in Clinical Cancer Research
  • A Four-Chemokine Signature Is Associated with a T-cell-Inflamed Phenotype in Primary and Metastatic Pancreatic Cancer, 2020, published in Clinical Cancer Research
  • Modification of BRCA1-associated breast cancer risk by HMMR overexpression, 2022, published in Nature Communications

Frequent coauthors collaborating with Khokha include Barbara T. Grünwald, Curtis W. McCloskey, Kazeera Aliar, Thomas Kislinger, and Foram Vyas.

Khokha's publications appear regularly in certain venues, indicating a focus within those research communities. Frequent publication venues include:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Cancer Research
  • Clinical Cancer Research
  • Nature Communications
  • British Journal of Cancer

The primary fields of study represented in Khokha's work cover Biochemistry, Genetics and Molecular Biology as well as Medicine. Subfields include:

  • Molecular Biology
  • Oncology
  • Cancer Research
  • Genetics
  • Immunology

Key research topics explored through Khokha's work are diverse and interconnected, focusing especially on cancer mechanisms and related cellular processes. These topics include:

  • Pancreatic and Hepatic Oncology Research
  • Epigenetics and DNA Methylation
  • Cancer Cells and Metastasis
  • Cancer Genomics and Diagnostics
  • Ubiquitin and proteasome pathways
  • Immune cells in cancer
  • Single-cell and spatial transcriptomics

Khokha has been recognized with awards including the Robert L. Noble Prize from the Canadian Cancer Society in 2014 and election as a Fellow of the Royal Society of Canada, Academy of Science, in 2020.

Best Publications

  • Regulation of cancer cell migration and bone metastasis by RANKL

    D. Holstead Jones;Tomoki Nakashima;Otto H. Sanchez;Otto H. Sanchez;Ivona Kozieradzki;Ivona Kozieradzki

  • The osteoclast differentiation factor osteoprotegerin-ligand is essential for mammary gland development

    Jimmie E Fata;Young-Yun Kong;Young-Yun Kong;Ji Li;Takehiko Sasaki

  • Glutathione and Thioredoxin Antioxidant Pathways Synergize to Drive Cancer Initiation and Progression

    Isaac S. Harris;Aislinn E. Treloar;Satoshi Inoue;Masato Sasaki

  • Progesterone induces adult mammary stem cell expansion

    Purna A. Joshi;Hartland W. Jackson;Alexander G. Beristain;Marco A. Di Grappa

  • Osteoclast differentiation factor RANKL controls development of progestin-driven mammary cancer.

    Daniel Schramek;Andreas Leibbrandt;Verena Sigl;Lukas Kenner

  • Suppression of tumor growth and metastasis in Mgat5-deficient mice.

    Maria Granovsky;Maria Granovsky;Jimmie Fata;Judy Pawling;William J. Muller

  • Tissue inhibitor of metalloproteinases-3 (TIMP-3) is an extracellular matrix-associated protein with a distinctive pattern of expression in mouse cells and tissues.

    K J Leco;R Khokha;N Pavloff;S P Hawkes

  • Metalloproteinases and their natural inhibitors in inflammation and immunity

    Rama Khokha;Aditya Murthy;Ashley Weiss

  • Antisense RNA-induced reduction in murine TIMP levels confers oncogenicity on Swiss 3T3 cells.

    Rama Khokha;Paul Waterhouse;Simcha Yagel;Peeyush K. Lala

  • Tumor Necrosis Factor-α Mediates Cardiac Remodeling and Ventricular Dysfunction After Pressure Overload State

    Mei Sun;Manyin Chen;Fayez Dawood;Urszula Zurawska

  • The Association of Measured Breast Tissue Characteristics with Mammographic Density and Other Risk Factors for Breast Cancer

    Tong Li;Limei Sun;Naomi Miller;Trudey Nicklee

  • TIMPs: versatile extracellular regulators in cancer

    Hartland W. Jackson;Hartland W. Jackson;Virginie Defamie;Paul Waterhouse;Rama Khokha

  • RhoC is dispensable for embryogenesis and tumor initiation but essential for metastasis

    Anne Hakem;Otto Sanchez-Sweatman;Annick You-Ten;Gordon Duncan

  • iRhom2 regulation of TACE controls TNF-mediated protection against Listeria and responses to LPS.

    David R. McIlwain;Philipp A. Lang;Philipp A. Lang;Thorsten Maretzky;Koichi Hamada

  • Molecular mechanisms of tumor invasion and metastasis: an integrated view.

    R. A. Cairns;R. Khokha;R. P. Hill

  • Steps in tumor metastasis: new concepts from intravital videomicroscopy.

    Ann F. Chambers;Ian C. MacDonald;Eric E. Schmidt;Sahadia Koop

  • Binding to EGF receptor of a laminin-5 EGF-like fragment liberated during MMP-dependent mammary gland involution

    Susann Schenk;Edith Hintermann;Martin Bilban;Naohiko Koshikawa

  • Abnormal TNF activity in Timp3 −/− mice leads to chronic hepatic inflammation and failure of liver regeneration

    Fazilat F Mohammed;David S Smookler;Suzanne E M Taylor;Barbara Fingleton

  • Spontaneous air space enlargement in the lungs of mice lacking tissue inhibitor of metalloproteinases-3 (TIMP-3)

    Kevin J. Leco;Paul Waterhouse;Otto H. Sanchez;Katrina L.M. Gowing

  • Matrix metalloproteinases and their tissue inhibitors direct cell fate during cancer development.

    C V Hojilla;F F Mohammed;R Khokha

Frequent Co-Authors

Josef M. Penninger
Josef M. Penninger University of British Columbia
Thomas Kislinger
Thomas Kislinger University of Toronto
Tak W. Mak
Tak W. Mak Princess Margaret Cancer Centre
Dylan R. Edwards
Dylan R. Edwards University of East Anglia
Carl P. Blobel
Carl P. Blobel Cornell University
Ann F. Chambers
Ann F. Chambers University of Western Ontario
Paul C. Boutros
Paul C. Boutros University of California, Los Angeles
Andrew Wakeham
Andrew Wakeham Princess Margaret Cancer Centre
David T. Denhardt
David T. Denhardt Rutgers, The State University of New Jersey
Tak W. Mak
Tak W. Mak Princess Margaret Cancer Centre

External Links

Personal website of Rama Khokha

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Each of these online degrees aligns with a strong foundation in Biology or Biochemistry, helping shape the skills and knowledge needed for advancement in the evolving healthcare field.

Best Scientists Citing Rama Khokha

Josef M. Penninger

Josef M. Penninger

University of British Columbia

Publications: 97

Gavin Y. Oudit

Gavin Y. Oudit

University of Alberta

Publications: 68

James W. Dennis

James W. Dennis

Lunenfeld-Tanenbaum Research Institute

Publications: 66

Ann F. Chambers

Ann F. Chambers

University of Western Ontario

Publications: 62

Carl P. Blobel

Carl P. Blobel

Cornell University

Publications: 57

Naoyuki Taniguchi

Naoyuki Taniguchi

Osaka University

Publications: 55

Dominique Heymann

Dominique Heymann

University of Nantes

Publications: 51

Gillian Murphy

Gillian Murphy

University of Cambridge

Publications: 49

Dylan R. Edwards

Dylan R. Edwards

University of East Anglia

Publications: 47

Zena Werb

Zena Werb

University of California, San Francisco

Publications: 46

William G. Stetler-Stevenson

William G. Stetler-Stevenson

National Institutes of Health

Publications: 44

Hideaki Nagase

Hideaki Nagase

University of Oxford

Publications: 41

Tak W. Mak

Tak W. Mak

Princess Margaret Cancer Centre

Publications: 38

Stefan Rose-John

Stefan Rose-John

Kiel University

Publications: 34

Geoffrey J. Lindeman

Geoffrey J. Lindeman

Peter MacCallum Cancer Centre

Publications: 32

Massimo Federici

Massimo Federici

University of Rome Tor Vergata

Publications: 32

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