World's Best Scientists 2026 revealed!

D-Index & Metrics

Biology and Biochemistry

D-Index
92
Citations
27267
World Ranking
2237
National Ranking
62

Medicine

D-Index
92
Citations
27438
World Ranking
11415
National Ranking
458

Rama Khokha publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Rama Khokha sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 199 publications — 49th percentile

49% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Rama Khokha D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Rama Khokha sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 92 D-Index — 89th percentile

89% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Research.com Recognitions

  • 2020 - Fellow of the Royal Society of Canada Academy of Science
  • 2014 - Robert L. Noble Prize, Canadian Cancer Society

Overview

Rama Khokha is affiliated with the Princess Margaret Cancer Centre in Canada and has contributed extensively to research in oncology, molecular biology, and medicine. Their work focuses on cancer biology, particularly pancreatic and hepatic oncology, cancer genomics, epigenetics, and immunology.

The scientist's recent publications illustrate key themes across tumor microenvironments, immune regulation, and cancer subtype differentiation. Notable papers include:

  • Tryptophan-derived microbial metabolites activate the aryl hydrocarbon receptor in tumor-associated macrophages to suppress anti-tumor immunity, 2022, published in Immunity
  • Spatially confined sub-tumor microenvironments in pancreatic cancer, 2021, published in Cell
  • GATA6 Expression Distinguishes Classical and Basal-like Subtypes in Advanced Pancreatic Cancer, 2020, published in Clinical Cancer Research
  • A Four-Chemokine Signature Is Associated with a T-cell-Inflamed Phenotype in Primary and Metastatic Pancreatic Cancer, 2020, published in Clinical Cancer Research
  • Modification of BRCA1-associated breast cancer risk by HMMR overexpression, 2022, published in Nature Communications

Frequent coauthors collaborating with Khokha include Barbara T. Grünwald, Curtis W. McCloskey, Kazeera Aliar, Thomas Kislinger, and Foram Vyas.

Khokha's publications appear regularly in certain venues, indicating a focus within those research communities. Frequent publication venues include:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Cancer Research
  • Clinical Cancer Research
  • Nature Communications
  • British Journal of Cancer

The primary fields of study represented in Khokha's work cover Biochemistry, Genetics and Molecular Biology as well as Medicine. Subfields include:

  • Molecular Biology
  • Oncology
  • Cancer Research
  • Genetics
  • Immunology

Key research topics explored through Khokha's work are diverse and interconnected, focusing especially on cancer mechanisms and related cellular processes. These topics include:

  • Pancreatic and Hepatic Oncology Research
  • Epigenetics and DNA Methylation
  • Cancer Cells and Metastasis
  • Cancer Genomics and Diagnostics
  • Ubiquitin and proteasome pathways
  • Immune cells in cancer
  • Single-cell and spatial transcriptomics

Khokha has been recognized with awards including the Robert L. Noble Prize from the Canadian Cancer Society in 2014 and election as a Fellow of the Royal Society of Canada, Academy of Science, in 2020.

Best Publications

  • Regulation of cancer cell migration and bone metastasis by RANKL

    D. Holstead Jones;Tomoki Nakashima;Otto H. Sanchez;Otto H. Sanchez;Ivona Kozieradzki;Ivona Kozieradzki

  • The osteoclast differentiation factor osteoprotegerin-ligand is essential for mammary gland development

    Jimmie E Fata;Young-Yun Kong;Young-Yun Kong;Ji Li;Takehiko Sasaki

  • Glutathione and Thioredoxin Antioxidant Pathways Synergize to Drive Cancer Initiation and Progression

    Isaac S. Harris;Aislinn E. Treloar;Satoshi Inoue;Masato Sasaki

  • Progesterone induces adult mammary stem cell expansion

    Purna A. Joshi;Hartland W. Jackson;Alexander G. Beristain;Marco A. Di Grappa

  • Osteoclast differentiation factor RANKL controls development of progestin-driven mammary cancer.

    Daniel Schramek;Andreas Leibbrandt;Verena Sigl;Lukas Kenner

  • Suppression of tumor growth and metastasis in Mgat5-deficient mice.

    Maria Granovsky;Maria Granovsky;Jimmie Fata;Judy Pawling;William J. Muller

  • Tissue inhibitor of metalloproteinases-3 (TIMP-3) is an extracellular matrix-associated protein with a distinctive pattern of expression in mouse cells and tissues.

    K J Leco;R Khokha;N Pavloff;S P Hawkes

  • Metalloproteinases and their natural inhibitors in inflammation and immunity

    Rama Khokha;Aditya Murthy;Ashley Weiss

  • Antisense RNA-induced reduction in murine TIMP levels confers oncogenicity on Swiss 3T3 cells.

    Rama Khokha;Paul Waterhouse;Simcha Yagel;Peeyush K. Lala

  • Tumor Necrosis Factor-α Mediates Cardiac Remodeling and Ventricular Dysfunction After Pressure Overload State

    Mei Sun;Manyin Chen;Fayez Dawood;Urszula Zurawska

  • The Association of Measured Breast Tissue Characteristics with Mammographic Density and Other Risk Factors for Breast Cancer

    Tong Li;Limei Sun;Naomi Miller;Trudey Nicklee

  • TIMPs: versatile extracellular regulators in cancer

    Hartland W. Jackson;Hartland W. Jackson;Virginie Defamie;Paul Waterhouse;Rama Khokha

  • RhoC is dispensable for embryogenesis and tumor initiation but essential for metastasis

    Anne Hakem;Otto Sanchez-Sweatman;Annick You-Ten;Gordon Duncan

  • iRhom2 regulation of TACE controls TNF-mediated protection against Listeria and responses to LPS.

    David R. McIlwain;Philipp A. Lang;Philipp A. Lang;Thorsten Maretzky;Koichi Hamada

  • Molecular mechanisms of tumor invasion and metastasis: an integrated view.

    R. A. Cairns;R. Khokha;R. P. Hill

  • Steps in tumor metastasis: new concepts from intravital videomicroscopy.

    Ann F. Chambers;Ian C. MacDonald;Eric E. Schmidt;Sahadia Koop

  • Binding to EGF receptor of a laminin-5 EGF-like fragment liberated during MMP-dependent mammary gland involution

    Susann Schenk;Edith Hintermann;Martin Bilban;Naohiko Koshikawa

  • Abnormal TNF activity in Timp3 −/− mice leads to chronic hepatic inflammation and failure of liver regeneration

    Fazilat F Mohammed;David S Smookler;Suzanne E M Taylor;Barbara Fingleton

  • Spontaneous air space enlargement in the lungs of mice lacking tissue inhibitor of metalloproteinases-3 (TIMP-3)

    Kevin J. Leco;Paul Waterhouse;Otto H. Sanchez;Katrina L.M. Gowing

  • Matrix metalloproteinases and their tissue inhibitors direct cell fate during cancer development.

    C V Hojilla;F F Mohammed;R Khokha

Frequent Co-Authors

Josef M. Penninger
Josef M. Penninger University of British Columbia
Thomas Kislinger
Thomas Kislinger University of Toronto
Tak W. Mak
Tak W. Mak Princess Margaret Cancer Centre
Dylan R. Edwards
Dylan R. Edwards University of East Anglia
Carl P. Blobel
Carl P. Blobel Cornell University
Ann F. Chambers
Ann F. Chambers University of Western Ontario
Paul C. Boutros
Paul C. Boutros University of California, Los Angeles
Andrew Wakeham
Andrew Wakeham Princess Margaret Cancer Centre
David T. Denhardt
David T. Denhardt Rutgers, The State University of New Jersey
Tak W. Mak
Tak W. Mak Princess Margaret Cancer Centre

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