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Timothy L. Denning

D-Index & Metrics

Discipline name D-Index World Ranking Current World Ranking National Ranking Current National Ranking Publications Citations
Immunology 48 4427 4094 2010 1843 117 9217

Timothy L. Denning publications per year

The chart shows the history of publications by Timothy L. Denning between 1996 and 2025, highlighting the no. of papers published in each year and offering an overview of the publication velocity of this scholar. Timothy L. Denning published across 30 years, from 1996 to 2025, averaging 4.9 papers a year. Output peaked at 12 publications in 2017. 5 of the 147 publications appeared in the last two years.

No. of publications
5 10
Bar chart. Horizontal axis: year, 1996 to 2025. Vertical axis: number of publications, 0 to 12. Peak 12 publications in 2017. 1996: 1 publication 1997: 0 publications 1998: 1 publication 1999: 2 publications 2000: 6 publications 2001: 4 publications 2002: 4 publications 2003: 5 publications 2004: 2 publications 2005: 1 publication 2006: 2 publications 2007: 3 publications 2008: 2 publications 2009: 2 publications 2010: 5 publications 2011: 8 publications 2012: 11 publications 2013: 8 publications 2014: 8 publications 2015: 10 publications 2016: 7 publications 2017: 12 publications 2018: 10 publications 2019: 7 publications 2020: 7 publications 2021: 5 publications 2022: 3 publications 2023: 6 publications 2024: 4 publications 2025: 1 publication
1996 2025

147 publications in total across all disciplines

View publications per year as a table
Timothy L. Denning: publications per year, 1996 to 2025
Year Publications
1996 1
1997 0
1998 1
1999 2
2000 6
2001 4
2002 4
2003 5
2004 2
2005 1
2006 2
2007 3
2008 2
2009 2
2010 5
2011 8
2012 11
2013 8
2014 8
2015 10
2016 7
2017 12
2018 10
2019 7
2020 7
2021 5
2022 3
2023 6
2024 4
2025 1
Total 147
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Timothy L. Denning publication distribution in Immunology in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Immunology in 2026. The highlighted bar marks where Timothy L. Denning sits on this spectrum.

No. of scientists
50 100 150
Bar chart with 76 bars. Horizontal axis: publications, 62–71 to 807+. Vertical axis: number of scientists, 0 to 189. Most scientists, 189, have 152–161 publications. The last bar groups every scientist with 807 publications or more. The highlighted bar, 112–121 publications, is where this scientist sits. 62–71 publications: 9 scientists 72–81 publications: 23 scientists 82–91 publications: 40 scientists 92–101 publications: 72 scientists 102–111 publications: 86 scientists 112–121 publications: 108 scientists 122–131 publications: 150 scientists 132–141 publications: 160 scientists 142–151 publications: 159 scientists 152–161 publications: 189 scientists 162–171 publications: 166 scientists 172–181 publications: 181 scientists 182–191 publications: 188 scientists 192–201 publications: 187 scientists 202–211 publications: 178 scientists 212–221 publications: 158 scientists 222–231 publications: 168 scientists 232–241 publications: 159 scientists 242–251 publications: 160 scientists 252–261 publications: 148 scientists 262–271 publications: 116 scientists 272–281 publications: 125 scientists 282–291 publications: 115 scientists 292–301 publications: 111 scientists 302–311 publications: 95 scientists 312–321 publications: 97 scientists 322–331 publications: 97 scientists 332–341 publications: 99 scientists 342–351 publications: 96 scientists 352–361 publications: 69 scientists 362–371 publications: 76 scientists 372–381 publications: 71 scientists 382–391 publications: 71 scientists 392–401 publications: 62 scientists 402–411 publications: 54 scientists 412–421 publications: 41 scientists 422–431 publications: 63 scientists 432–441 publications: 53 scientists 442–451 publications: 31 scientists 452–461 publications: 42 scientists 462–471 publications: 40 scientists 472–481 publications: 29 scientists 482–491 publications: 34 scientists 492–501 publications: 30 scientists 502–511 publications: 23 scientists 512–521 publications: 41 scientists 522–531 publications: 29 scientists 532–541 publications: 28 scientists 542–551 publications: 16 scientists 552–561 publications: 18 scientists 562–571 publications: 18 scientists 572–581 publications: 17 scientists 582–591 publications: 17 scientists 592–601 publications: 17 scientists 602–611 publications: 19 scientists 612–621 publications: 13 scientists 622–631 publications: 12 scientists 632–641 publications: 11 scientists 642–651 publications: 16 scientists 652–661 publications: 9 scientists 662–671 publications: 5 scientists 672–681 publications: 13 scientists 682–691 publications: 12 scientists 692–701 publications: 6 scientists 702–711 publications: 6 scientists 712–721 publications: 9 scientists 722–731 publications: 9 scientists 732–741 publications: 6 scientists 742–751 publications: 11 scientists 752–761 publications: 10 scientists 762–771 publications: 7 scientists 772–781 publications: 12 scientists 782–791 publications: 7 scientists 792–801 publications: 5 scientists 802–806 publications: 1 scientist 807+ publications: 100 scientists
62–71 publications 807+

This scientist: 117 publications — 6th percentile

6% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 807 publications or more.

View publications distribution as a table
Number of Immunology scientists by publication count, Research.com 2026 ranking edition. Based on 4,929 ranked scientists.
Publications Scientists This scientist
62–71 9
72–81 23
82–91 40
92–101 72
102–111 86
112–121 108 117
122–131 150
132–141 160
142–151 159
152–161 189
162–171 166
172–181 181
182–191 188
192–201 187
202–211 178
212–221 158
222–231 168
232–241 159
242–251 160
252–261 148
262–271 116
272–281 125
282–291 115
292–301 111
302–311 95
312–321 97
322–331 97
332–341 99
342–351 96
352–361 69
362–371 76
372–381 71
382–391 71
392–401 62
402–411 54
412–421 41
422–431 63
432–441 53
442–451 31
452–461 42
462–471 40
472–481 29
482–491 34
492–501 30
502–511 23
512–521 41
522–531 29
532–541 28
542–551 16
552–561 18
562–571 18
572–581 17
582–591 17
592–601 17
602–611 19
612–621 13
622–631 12
632–641 11
642–651 16
652–661 9
662–671 5
672–681 13
682–691 12
692–701 6
702–711 6
712–721 9
722–731 9
732–741 6
742–751 11
752–761 10
762–771 7
772–781 12
782–791 7
792–801 5
802–806 1
807+ 100
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Timothy L. Denning D-index placement in Immunology in 2026

The chart shows the D-index (discipline H-index) distribution of Immunology scientists ranked by Research.com in 2026. The highlighted bar marks where Timothy L. Denning sits on this spectrum.

No. of scientists
50 100 150 200
Bar chart with 60 bars. Horizontal axis: D-Index, 40–41 to 157+. Vertical axis: number of scientists, 0 to 201. Most scientists, 201, have 58–59 D-Index. The last bar groups every scientist with 157 D-Index or more. The highlighted bar, 48–49 D-Index, is where this scientist sits. 40–41 D-Index: 28 scientists 42–43 D-Index: 91 scientists 44–45 D-Index: 157 scientists 46–47 D-Index: 192 scientists 48–49 D-Index: 175 scientists 50–51 D-Index: 188 scientists 52–53 D-Index: 177 scientists 54–55 D-Index: 155 scientists 56–57 D-Index: 194 scientists 58–59 D-Index: 201 scientists 60–61 D-Index: 197 scientists 62–63 D-Index: 201 scientists 64–65 D-Index: 173 scientists 66–67 D-Index: 167 scientists 68–69 D-Index: 172 scientists 70–71 D-Index: 199 scientists 72–73 D-Index: 184 scientists 74–75 D-Index: 133 scientists 76–77 D-Index: 163 scientists 78–79 D-Index: 127 scientists 80–81 D-Index: 121 scientists 82–83 D-Index: 125 scientists 84–85 D-Index: 121 scientists 86–87 D-Index: 102 scientists 88–89 D-Index: 96 scientists 90–91 D-Index: 75 scientists 92–93 D-Index: 72 scientists 94–95 D-Index: 74 scientists 96–97 D-Index: 66 scientists 98–99 D-Index: 68 scientists 100–101 D-Index: 49 scientists 102–103 D-Index: 60 scientists 104–105 D-Index: 54 scientists 106–107 D-Index: 36 scientists 108–109 D-Index: 23 scientists 110–111 D-Index: 52 scientists 112–113 D-Index: 41 scientists 114–115 D-Index: 34 scientists 116–117 D-Index: 25 scientists 118–119 D-Index: 28 scientists 120–121 D-Index: 11 scientists 122–123 D-Index: 20 scientists 124–125 D-Index: 27 scientists 126–127 D-Index: 19 scientists 128–129 D-Index: 20 scientists 130–131 D-Index: 16 scientists 132–133 D-Index: 17 scientists 134–135 D-Index: 14 scientists 136–137 D-Index: 15 scientists 138–139 D-Index: 9 scientists 140–141 D-Index: 11 scientists 142–143 D-Index: 9 scientists 144–145 D-Index: 9 scientists 146–147 D-Index: 7 scientists 148–149 D-Index: 7 scientists 150–151 D-Index: 7 scientists 152–153 D-Index: 5 scientists 154–155 D-Index: 8 scientists 156 D-Index: 6 scientists 157+ D-Index: 96 scientists
40–41 D-Index 157+

This scientist: 48 D-Index — 11th percentile

11% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 157 D-Index or more.

View D-Index distribution as a table
Number of Immunology scientists by D-index, Research.com 2026 ranking edition. Based on 4,929 ranked scientists.
D-Index Scientists This scientist
40–41 28
42–43 91
44–45 157
46–47 192
48–49 175 48
50–51 188
52–53 177
54–55 155
56–57 194
58–59 201
60–61 197
62–63 201
64–65 173
66–67 167
68–69 172
70–71 199
72–73 184
74–75 133
76–77 163
78–79 127
80–81 121
82–83 125
84–85 121
86–87 102
88–89 96
90–91 75
92–93 72
94–95 74
96–97 66
98–99 68
100–101 49
102–103 60
104–105 54
106–107 36
108–109 23
110–111 52
112–113 41
114–115 34
116–117 25
118–119 28
120–121 11
122–123 20
124–125 27
126–127 19
128–129 20
130–131 16
132–133 17
134–135 14
136–137 15
138–139 9
140–141 11
142–143 9
144–145 9
146–147 7
148–149 7
150–151 7
152–153 5
154–155 8
156 6
157+ 96
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Overview

Timothy L. Denning is affiliated with the Georgia Research Alliance in the United States and has a research focus primarily in the fields of Immunology and Microbiology, Medicine, and Biochemistry, Genetics and Molecular Biology. Their scientific work spans multiple subfields, including Immunology, Molecular Biology, Epidemiology, Genetics, and Dermatology.

Their research addresses a variety of topics, with notable coverage in:

  • Psoriasis: Treatment and Pathogenesis
  • Epigenetics and DNA Methylation
  • Immune Cell Function and Interaction
  • Immune Response and Inflammation
  • Gut microbiota and health
  • Dermatology and Skin Diseases
  • Immunotherapy and Immune Responses

Denning's published works include articles in several frequent venues, including:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Nature Communications
  • Immunology
  • ACS Omega
  • Mucosal Immunology

Some of the recent publications by Timothy L. Denning include:

  • "PTH induces bone loss via microbial-dependent expansion of intestinal TNF+ T cells and Th17 cells", 2020, Nature Communications
  • "Internalization of Garlic-Derived Nanovesicles on Liver Cells is Triggered by Interaction With CD98", 2020, ACS Omega
  • "IL-36 cytokines and gut immunity", 2021, Immunology
  • "Erythroid differentiation regulator-1 induced by microbiota in early life drives intestinal stem cell proliferation and regeneration", 2020, Nature Communications
  • "Self and microbiota-derived epitopes induce CD4+ T cell anergy and conversion into CD4+Foxp3+ regulatory cells", 2020, Mucosal Immunology

The scientist collaborates frequently with several co-authors, including:

  • Vu L. Ngo
  • Andrew T. Gewirtz
  • Michal Kuczma
  • Hirohito Abo
  • Yuning Hou

Best Publications

  • Lamina propria macrophages and dendritic cells differentially induce regulatory and interleukin 17-producing T cell responses.

    Timothy L Denning;Yi-chong Wang;Seema R Patel;Ifor R Williams

  • Yeast zymosan, a stimulus for TLR2 and dectin-1, induces regulatory antigen-presenting cells and immunological tolerance

    Stephanie Dillon;Sudhanshu Agrawal;Kaustuv Banerjee;John Letterio

  • Functional Specializations of Intestinal Dendritic Cell and Macrophage Subsets That Control Th17 and Regulatory T Cell Responses Are Dependent on the T Cell/APC Ratio, Source of Mouse Strain, and Regional Localization

    Timothy L. Denning;Brian A. Norris;Oscar Medina-Contreras;Santhakumar Manicassamy

  • Thymus-derived regulatory T cells contribute to tolerance to commensal microbiota

    Anna Cebula;Michal Seweryn;Michal Seweryn;Grzegorz A. Rempala;Simarjot Singh Pabla

  • Toll-like receptor 2-dependent induction of vitamin A-metabolizing enzymes in dendritic cells promotes T regulatory responses and inhibits autoimmunity.

    Santhakumar Manicassamy;Rajesh Ravindran;Jiusheng Deng;Herold Oluoch

  • CX3CR1 regulates intestinal macrophage homeostasis, bacterial translocation and colitogenic TH17 responses in mice

    Oscar Medina-Contreras;Duke Geem;Oskar Laur;Ifor R Williams

  • CX3CR1 regulates intestinal macrophage homeostasis, bacterial translocation, and colitogenic Th17 responses in mice

    Oscar Medina-Contreras;Duke Geem;Oskar Laur;Ifor R. Williams

  • Wound repair: role of immune–epithelial interactions

    G. Leoni;P. A. Neumann;R. Sumagin;T. L. Denning

  • Prevention and cure of rotavirus infection via TLR5/NLRC4–mediated production of IL-22 and IL-18

    Benyue Zhang;Benoit Chassaing;Zhenda Shi;Robin Uchiyama

  • Macrophage-derived IL-10 mediates mucosal repair by epithelial WISP-1 signaling

    Miguel Quiros;Hikaru Nishio;Philipp A Neumann;Dorothee Siuda

  • Division of labor, plasticity, and crosstalk between dendritic cell subsets

    Bali Pulendran;Hua Tang;Timothy L. Denning

  • ERK1-/- mice exhibit Th1 cell polarization and increased susceptibility to experimental autoimmune encephalomyelitis.

    Anshu Agrawal;Stephanie Dillon;Timothy L. Denning;Bali Pulendran

  • Mouse TCRαβ+CD8αα Intraepithelial Lymphocytes Express Genes That Down-Regulate Their Antigen Reactivity and Suppress Immune Responses

    Timothy L. Denning;Steve Granger;Daniel Mucida;Ryan Graddy

  • Pulmonary antigen encounter regulates the establishment of tissue-resident CD8 memory T cells in the lung airways and parenchyma.

    Sean R. McMaster;Alexander N. Wein;Paul R. Dunbar;Sarah L. Hayward

  • Differential Induction of Interleukin-10 and Interleukin-12 in Dendritic Cells by Microbial Toll-Like Receptor Activators and Skewing of T-Cell Cytokine Profiles

    Hai Qi;Timothy L. Denning;Lynn Soong

  • Oxidative stress induces the expression of Fas and Fas ligand and apoptosis in murine intestinal epithelial cells.

    Timothy L Denning;Hiromasa Takaishi;Sheila E Crowe;Istvan Boldogh

  • Fab’-bearing siRNA TNFα-loaded nanoparticles targeted to colonic macrophages offer an effective therapy for experimental colitis

    Hamed Laroui;Emilie Viennois;Bo Xiao;Brandon S.B. Canup

  • Cutting Edge: CD4+CD25+ Regulatory T Cells Impaired for Intestinal Homing Can Prevent Colitis

    Timothy L. Denning;Gisen Kim;Mitchell Kronenberg

  • Hepatic Stellate Cells Preferentially Induce Foxp3+ Regulatory T Cells by Production of Retinoic Acid

    Richard M. Dunham;Manoj Thapa;Victoria M. Velazquez;Elizabeth J. Elrod

  • Negative selection of T cells by Helicobacter pylori as a model for bacterial strain selection by immune evasion.

    Jide Wang;Edward G. Brooks;Kathleen B. Bamford;Timothy L. Denning

  • IL-17A-mediated neutrophil recruitment limits expansion of segmented filamentous bacteria

    K L Flannigan;K L Flannigan;V L Ngo;D Geem;D Geem;A Harusato

  • Cutting Edge: IL-36 Receptor Promotes Resolution of Intestinal Damage.

    Oscar Medina-Contreras;Akihito Harusato;Hikaru Nishio;Kyle L. Flannigan

  • Compromised Intestinal Epithelial Barrier Induces Adaptive Immune Compensation that Protects from Colitis

    Manirath Khounlotham;Wooki Kim;Eric Peatman;Porfirio Nava

Frequent Co-Authors

Asma Nusrat
Asma Nusrat University of Michigan–Ann Arbor
Charles A. Parkos
Charles A. Parkos University of Michigan–Ann Arbor
Andrew T. Gewirtz
Andrew T. Gewirtz Georgia State University
Didier Merlin
Didier Merlin Georgia State University
Peter B. Ernst
Peter B. Ernst University of California, San Diego
Bali Pulendran
Bali Pulendran Stanford University
Benoit Chassaing
Benoit Chassaing Georgia State University
Ifor R. Williams
Ifor R. Williams Emory University
Shanthi V. Sitaraman
Shanthi V. Sitaraman Emory University
Sheila E. Crowe
Sheila E. Crowe University of California, San Diego

External Links

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Best Scientists Citing Timothy L. Denning

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Didier Merlin

Didier Merlin

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Asma Nusrat

Asma Nusrat

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Publications: 30

Peter B. Ernst

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Publications: 26

Nobuhiko Kamada

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Publications: 26

Bali Pulendran

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Stanford University

Publications: 25

Mihai G. Netea

Mihai G. Netea

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Publications: 24

Kenya Honda

Kenya Honda

Keio University

Publications: 22

Sheila E. Crowe

Sheila E. Crowe

University of California, San Diego

Publications: 21

Hilde Cheroutre

Hilde Cheroutre

La Jolla Institute For Allergy & Immunology

Publications: 20

Yasmine Belkaid

Yasmine Belkaid

National Institute of Allergy and Infectious Diseases

Publications: 20

Andrew T. Gewirtz

Andrew T. Gewirtz

Georgia State University

Publications: 20

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Jacques Banchereau

Immunai

Publications: 20

Bernard Malissen

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Aix-Marseille University

Publications: 19

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Publications: 19

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Kitasato University

Publications: 19

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