World's Best Scientists 2026 revealed!
VIEW LIST
Tim Formosa

D-Index & Metrics

Biology and Biochemistry

D-Index
42
Citations
6075
World Ranking
19499
National Ranking
7950

Tim Formosa publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Tim Formosa sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 78 publications — 1st percentile

1% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Tim Formosa D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Tim Formosa sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 42 D-Index — 1st percentile

1% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Overview

Tim Formosa is affiliated with the University of Utah in the United States. Their research primarily focuses on the field of Biochemistry, Genetics and Molecular Biology, with a significant concentration in Molecular Biology. Their work spans various topics including Genomics and Chromatin Dynamics, RNA modifications and cancer, DNA Repair Mechanisms, RNA Research and Splicing, RNA and protein synthesis mechanisms, Innovations in Medical Education, and Ubiquitin and proteasome pathways.

Key recent publications by Tim Formosa include:

  • The role of FACT in managing chromatin: disruption, assembly, or repair? (2020), Nucleic Acids Research
  • Electron microscopy analysis of ATP-independent nucleosome unfolding by FACT (2022), Communications Biology
  • The interaction between the Spt6-tSH2 domain and Rpb1 affects multiple functions of RNA Polymerase II (2021), Nucleic Acids Research
  • Hmo1 Protein Affects the Nucleosome Structure and Supports the Nucleosome Reorganization Activity of Yeast FACT (2022), Cells
  • Mechanism of curaxin-dependent nucleosome unfolding by FACT (2022), Frontiers in Molecular Biosciences

Frequent co-authors collaborating with Tim Formosa include:

  • Anastasiia L. Sivkina
  • М. П. Кирпичников
  • Vasily M. Studitsky
  • Maria E. Valieva
  • Laura McCullough

The scientist has published repeatedly in several key venues, such as:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Nucleic Acids Research
  • Communications Biology
  • Cells
  • Frontiers in Molecular Biosciences

Tim Formosa's considerable contributions span the study of chromatin dynamics and nucleosome behavior, RNA processes, and mechanisms involved in DNA repair and protein synthesis. Their collaborations and publications highlight a sustained research interest in understanding molecular mechanisms related to gene expression and chromatin regulation.

Best Publications

  • DNA synthesis dependent on genetic recombination: Characterization of a reaction catalyzed by purified bacteriophage T4 proteins

    Timothy Formosa;Bruce M. Alberts

  • Spt16–Pob3 and the HMG protein Nhp6 combine to form the nucleosome‐binding factor SPN

    Tim Formosa;Peter Eriksson;Jacqui Wittmeyer;Jennifer Ginn

  • Studies on DNA replication in the bacteriophage T4 in vitro system.

    B. M. Alberts;J. Barry;P. Bedinger;T. Formosa

  • The Saccharomyces cerevisiae DNA polymerase alpha catalytic subunit interacts with Cdc68/Spt16 and with Pob3, a protein similar to an HMG1-like protein.

    Jacqueline Wittmeyer;Tim Formosa

  • The Role of FACT in Making and Breaking Nucleosomes

    Tim Formosa

  • Defects in SPT16 or POB3 (yFACT) in saccharomyces cerevisiae cause dependence on the Hir/Hpc pathway: Polymerase passage may degrade chromatin structure

    Tim Formosa;Susan Ruone;Melissa D. Adams;Aileen E. Olsen

  • yFACT Induces Global Accessibility of Nucleosomal DNA without H2A-H2B Displacement

    Hua Xin;Shinya Takahata;Mary Blanksma;Laura McCullough

  • Affinity purification of bacteriophage T4 proteins essential for DNA replication and genetic recombination.

    Timothy Formosa;Rae Lyn Burke;Bruce M. Alberts

  • Purification and characterization of the T4 bacteriophage uvsX protein.

    T Formosa;B M Alberts

  • Spt16 and Pob3 of Saccharomyces cerevisiae Form an Essential, Abundant Heterodimer That Is Nuclear, Chromatin-Associated, and Copurifies with DNA Polymerase α

    Jacqueline Wittmeyer;Lisa Joss;Tim Formosa

  • Structure of a Blm10 Complex Reveals Common Mechanisms for Proteasome Binding and Gate Opening

    Kianoush Sadre-Bazzaz;Frank G. Whitby;Howard Robinson;Tim Formosa

  • The Structure of the yFACT Pob3-M Domain, Its Interaction with the DNA Replication Factor RPA, and a Potential Role in Nucleosome Deposition

    Andrew P. VanDemark;Mary Blanksma;Elliott Ferris;Annie Heroux

  • Dna2 mutants reveal interactions with Dna polymerase alpha and Ctf4, a Pol alpha accessory factor, and show that full Dna2 helicase activity is not essential for growth.

    Tim Formosa;Thalia Nittis

  • FACT Disrupts Nucleosome Structure by Binding H2A-H2B with Conserved Peptide Motifs

    David J. Kemble;Laura L. McCullough;Frank G. Whitby;Tim Formosa

  • POB3 is required for both transcription and replication in the yeast Saccharomyces cerevisiae.

    Mylynda B. Schlesinger;Tim Formosa

  • Evidence that POB1, a Saccharomyces cerevisiae protein that binds to DNA polymerase alpha, acts in DNA metabolism in vivo.

    Jeff Miles;Tim Formosa

  • The yeast FACT complex has a role in transcriptional initiation.

    Debabrata Biswas;Yaxin Yu;Matthew Prall;Tim Formosa

  • Large-scale ATP-independent nucleosome unfolding by a histone chaperone

    Maria E Valieva;Grigoriy A Armeev;Kseniya S Kudryashova;Kseniya S Kudryashova;Nadezhda S Gerasimova

  • Multiple Nhp6 molecules are required to recruit Spt16-Pob3 to form yFACT complexes and to reorganize nucleosomes.

    Susan Ruone;Alison R. Rhoades;Tim Formosa

  • Structural and Functional Analysis of the Spt16p N-terminal Domain Reveals Overlapping Roles of yFACT Subunits

    Andrew P. VanDemark;Hua Xin;Laura McCullough;Robert Rawlins

Frequent Co-Authors

Christopher P. Hill
Christopher P. Hill University of Utah
David J. Stillman
David J. Stillman University of Utah
Bruce Alberts
Bruce Alberts University of California, San Francisco
Cynthia Wolberger
Cynthia Wolberger Johns Hopkins University School of Medicine
Ashraf Brik
Ashraf Brik Technion – Israel Institute of Technology
Howard Robinson
Howard Robinson Brookhaven National Laboratory
Stephen Buratowski
Stephen Buratowski Harvard University
Paul D. Kaufman
Paul D. Kaufman University of Massachusetts Chan Medical School
Danesh Moazed
Danesh Moazed Harvard University
Fred Winston
Fred Winston Harvard University

External Links

Personal website of Tim Formosa

If you think any of the details on this page are incorrect, let us know.

Report an issue

We appreciate your kind effort to assist us to improve this page, it would be helpful providing us with as much detail as possible in the text box below:

Related Online Degrees & Career Pathways

Pursuing a degree in Biology or Biochemistry opens doors to many related healthcare and science careers. Increasingly, students are leveraging flexible online options to earn credentials and advance in fields closely allied with life sciences.

For those considering nursing, online associate rn programs allow you to start a registered nursing career and gain clinical knowledge remotely, making it possible to balance education with work or family commitments.

Interested in preventive health or wellness? A nutritionist degree online can be a valuable path, blending scientific study with practical, client-facing skills for careers in healthcare, fitness, and education.

Students aspiring to leadership roles might explore top cahme-accredited mha programs, which emphasize healthcare management and administration—key in hospitals, labs, and biotech companies.

Meanwhile, those seeking advanced clinical positions can consider online dnp programs without dissertation, which streamline the process for earning a Doctor of Nursing Practice degree, often without the traditional research component.

Best Scientists Citing Tim Formosa

Karolin Luger

Karolin Luger

University of Colorado Boulder

Publications: 26

Stephen J. Benkovic

Stephen J. Benkovic

Pennsylvania State University

Publications: 23

Bruce Alberts

Bruce Alberts

University of California, San Francisco

Publications: 23

Mike O'Donnell

Mike O'Donnell

Rockefeller University

Publications: 22

Judith L. Campbell

Judith L. Campbell

California Institute of Technology

Publications: 22

Fred Winston

Fred Winston

Harvard University

Publications: 21

Kenneth N. Kreuzer

Kenneth N. Kreuzer

Duke University

Publications: 21

David J. Stillman

David J. Stillman

University of Utah

Publications: 17

Brian D. Strahl

Brian D. Strahl

University of North Carolina at Chapel Hill

Publications: 17

Thomas Kodadek

Thomas Kodadek

Scripps Research Institute

Publications: 16

Huilin Li

Huilin Li

Van Andel Institute

Publications: 15

Charles C. Richardson

Charles C. Richardson

Harvard University

Publications: 15

Danny Reinberg

Danny Reinberg

University of Miami

Publications: 15

James E. Haber

James E. Haber

Brandeis University

Publications: 14

Zhiguo Zhang

Zhiguo Zhang

Columbia University

Publications: 13

Karim Labib

Karim Labib

University of Dundee

Publications: 13

Trending Scientists

Junqiao Wu

Junqiao Wu

University of California, Berkeley

Shaun M. Eack

Shaun M. Eack

University of Pittsburgh

Steve M. Jex

Steve M. Jex

Bowling Green State University

Lihong V. Wang

Lihong V. Wang

California Institute of Technology

Jorge Blanco

Jorge Blanco

University of Santiago de Compostela

Marina A. Freudenberg

Marina A. Freudenberg

University of Freiburg

Mark D. Fricker

Mark D. Fricker

University of Oxford

Nicholas J. Clifford

Nicholas J. Clifford

Loughborough University

Kim D. Raine

Kim D. Raine

University of Alberta

Tak H. Lee

Tak H. Lee

King's College London

S. Iniyan

S. Iniyan

Anna University, Chennai

Leah Findlater

Leah Findlater

University of Washington

Massimo Mecella

Massimo Mecella

Sapienza University of Rome

M. T. Rodgers

M. T. Rodgers

Wayne State University

Rainer Pöttgen

Rainer Pöttgen

University of Münster

José Paulo Sousa

José Paulo Sousa

University of Coimbra

Related Biology and Biochemistry Rankings & Guides

2026 Different Types of Regulatory Science & Affairs Degrees and Their Salaries

2026 Different Types of Regulatory Science & Affairs Degrees and Their Salaries
2026 How to Become a Healthcare Risk Manager: Education, Salary, and Job Outlook

2026 How to Become a Healthcare Risk Manager: Education, Salary, and Job Outlook
2026 Can You Get Into a Public Health Degree Master's Program with a Low GPA? Admission Chances & Workarounds

2026 Can You Get Into a Public Health Degree Master's Program with a Low GPA? Admission Chances & Workarounds
2026 MRI Tech vs. Radiology Tech: Explaining the Difference

2026 MRI Tech vs. Radiology Tech: Explaining the Difference
2026 Nutrition Degree vs. Certificate: Which Should You Choose?

2026 Nutrition Degree vs. Certificate: Which Should You Choose?
2026 Biotechnology Degree vs. Certificate: Which Should You Choose?

2026 Biotechnology Degree vs. Certificate: Which Should You Choose?
2026 Can You Get Into a Psychiatric Mental Health Nurse Practitioner Program with a Low GPA? Admission Chances & Workarounds

2026 Can You Get Into a Psychiatric Mental Health Nurse Practitioner Program with a Low GPA? Admission Chances & Workarounds
2026 Military-Friendly Online Nutrition Degree Programs: Benefits, Accreditation, and Career Outcomes

2026 Military-Friendly Online Nutrition Degree Programs: Benefits, Accreditation, and Career Outcomes
2026 Can You Get a Biology Degree Master's Without a Related Bachelor's Degree?

2026 Can You Get a Biology Degree Master's Without a Related Bachelor's Degree?
2026 How Fast Can You Get a Sports Science Degree Online?

2026 How Fast Can You Get a Sports Science Degree Online?
2026 Public Health vs. Healthcare Management Degree: Explaining the Difference

2026 Public Health vs. Healthcare Management Degree: Explaining the Difference
2026 Cheapest and Shortest Online Ultrasound Technology Degree Programs That Pay Well: Cost, Duration, and Salary Outlook

2026 Cheapest and Shortest Online Ultrasound Technology Degree Programs That Pay Well: Cost, Duration, and Salary Outlook
2026 Cheapest and Shortest Online Occupational Therapy Degree Programs That Pay Well: Cost, Duration, and Salary Outlook

2026 Cheapest and Shortest Online Occupational Therapy Degree Programs That Pay Well: Cost, Duration, and Salary Outlook
2026 Can You Get Into a Family Nurse Practitioner Program with a Low GPA? Admission Chances & Workarounds

2026 Can You Get Into a Family Nurse Practitioner Program with a Low GPA? Admission Chances & Workarounds
2026 Best Speech Pathology Degrees for Working Adults

2026 Best Speech Pathology Degrees for Working Adults
2026 Is Health Information Management a Hard Major? What Students Should Know

2026 Is Health Information Management a Hard Major? What Students Should Know
2026 Nutrition Degree Programs for Career Changers

2026 Nutrition Degree Programs for Career Changers
2026 Careers for Biology Majors That Pay Well

2026 Careers for Biology Majors That Pay Well
2026 Cheapest Online Psychiatric Mental Health Nurse Practitioner Master's Degrees That Pay Well: Tuition, Duration, and Earnings

2026 Cheapest Online Psychiatric Mental Health Nurse Practitioner Master's Degrees That Pay Well: Tuition, Duration, and Earnings
2026 Cheapest and Shortest Online Health Informatics Degree Programs That Pay Well: Cost, Duration, and Salary Outlook

2026 Cheapest and Shortest Online Health Informatics Degree Programs That Pay Well: Cost, Duration, and Salary Outlook
2026 Fastest Online Nutrition Degree Programs You Can Finish in Under 2 Years

2026 Fastest Online Nutrition Degree Programs You Can Finish in Under 2 Years

Recently Published Articles

2026 Best AI Degrees for Students Interested in Health Informatics

2026 Best AI Degrees for Students Interested in Health Informatics
2026 Different Types of Security Management Degrees and Their Salaries

2026 Different Types of Security Management Degrees and Their Salaries
2026 Fastest Online Master’s in Legal Studies Program

2026 Fastest Online Master’s in Legal Studies Program
2026 Most Affordable Online Animation Degree Programs

2026 Most Affordable Online Animation Degree Programs
2026 Most Affordable Online Master’s in Substance Abuse Counseling Degrees

2026 Most Affordable Online Master’s in Substance Abuse Counseling Degrees
2026 How to Become a Nutritionist in Virginia

2026 How to Become a Nutritionist in Virginia
2026 How to Become a Licensed Therapist (LPC) in Omaha, NE

2026 How to Become a Licensed Therapist (LPC) in Omaha, NE
2026 Online Artificial Intelligence Degree Master's Programs with No GRE or GMAT Requirements

2026 Online Artificial Intelligence Degree Master's Programs with No GRE or GMAT Requirements
2026 Cheapest Online Colleges That Accept FAFSA

2026 Cheapest Online Colleges That Accept FAFSA
2026 Credit Requirements for a Forensic Science Degree Explained

2026 Credit Requirements for a Forensic Science Degree Explained
2026 Is a Psychology Degree Worth It?

2026 Is a Psychology Degree Worth It?
2026 Online Accounting Bachelor's Degree Programs That Accept Transfer Credits

2026 Online Accounting Bachelor's Degree Programs That Accept Transfer Credits
2026 Most Affordable Online BS in Organizational Leadership Degree Programs

2026 Most Affordable Online BS in Organizational Leadership Degree Programs
2026 Online Communications Bachelor's Degree Programs With Accelerated Courses

2026 Online Communications Bachelor's Degree Programs With Accelerated Courses
2026 How Fast Can You Get a Forensic Science Degree Online?

2026 How Fast Can You Get a Forensic Science Degree Online?
2026 Is Library Science a Hard Major? What Students Should Know

2026 Is Library Science a Hard Major? What Students Should Know
2026 How to Become an HVAC Tech in Texas

2026 How to Become an HVAC Tech in Texas
2026 Fastest Online MSN in Nursing Education Programs

2026 Fastest Online MSN in Nursing Education Programs
2026 How Fast Can You Get a Geography Degree Online?

2026 How Fast Can You Get a Geography Degree Online?
2026 AI, Automation, and the Future of Industrial Engineering Degree Careers

2026 AI, Automation, and the Future of Industrial Engineering Degree Careers
2026 Cheapest Online Accounting Bachelor's Degrees That Pay Well: Tuition, Duration, and Earnings

2026 Cheapest Online Accounting Bachelor's Degrees That Pay Well: Tuition, Duration, and Earnings