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Johanna Myllyharju

Johanna Myllyharju

University of Oulu
Finland

D-Index & Metrics

Biology and Biochemistry

D-Index
68
Citations
16782
World Ranking
7779
National Ranking
56

Johanna Myllyharju publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Johanna Myllyharju sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 413 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 848 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 949 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 840 scientists 207–216 publications: 733 scientists 217–226 publications: 708 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 417 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 196 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 59 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 175 publications — 39th percentile

39% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Johanna Myllyharju D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Johanna Myllyharju sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 316 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 899 scientists 52–53 D-Index: 1,025 scientists 54–55 D-Index: 1,149 scientists 56–57 D-Index: 1,235 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,162 scientists 62–63 D-Index: 1,130 scientists 64–65 D-Index: 1,031 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 714 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 71 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 68 D-Index — 61st percentile

61% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Overview

Johanna Myllyharju is affiliated with the University of Oulu in Finland and has contributed extensively to the fields of biochemistry, genetics, and molecular biology, with significant research output in medicine as well. Their work notably spans multiple subfields including molecular biology, cancer research, genetics, physiology, and pulmonary and respiratory medicine.

The scientist's research topics cover a range of areas primarily focused on cancer, hypoxia, and metabolism. Other main themes in their research include cancer-related gene regulation, high altitude and hypoxia, adipose tissue and metabolism, mitochondrial function and pathology, connective tissue disorders research, and ubiquitin and proteasome pathways.

Some of the frequent collaborators in Johanna Myllyharju's research include:

  • Peppi Koivunen
  • Antti M. Salo
  • Joni M. Mäki
  • Raisa Serpi
  • M. Kristian Koski

The scientist has published multiple papers in several reputable venues. The most common publication outlets for their work are:

  • Journal of Biological Chemistry
  • bioRxiv (Cold Spring Harbor Laboratory)
  • Matrix Biology
  • Experimental Dermatology
  • Angiogenesis

Representative recent publications highlight the breadth of Johanna Myllyharju's research, including:

  • Prolyl and lysyl hydroxylases in collagen synthesis, 2020, Experimental Dermatology
  • Type I and type V procollagen triple helix uses different subsets of the molecular ensemble for lysine posttranslational modifications in the rER, 2021, Journal of Biological Chemistry
  • PHD2 deletion in endothelial or arterial smooth muscle cells reveals vascular cell type-specific responses in pulmonary hypertension and fibrosis, 2022, Angiogenesis
  • Reduced Bone Mass in Collagen Prolyl 4-Hydroxylase P4ha1 +/−; P4ha2 −/− Compound Mutant Mice, 2022, JBMR Plus
  • HIF-P4H-2 inhibition enhances intestinal fructose metabolism and induces thermogenesis protecting against NAFLD, 2020, Journal of Molecular Medicine

Best Publications

  • Collagens, modifying enzymes and their mutations in humans, flies and worms

    Johanna Myllyharju;Kari I. Kivirikko

  • Collagens and collagen-related diseases

    Johanna Myllyharju;Kari I Kivirikko

  • Characterization of the human prolyl 4-hydroxylases that modify the hypoxia-inducible factor.

    Maija Hirsilä;Peppi Koivunen;Volkmar Günzler;Kari I. Kivirikko

  • Inhibition of Hypoxia-inducible Factor (HIF) Hydroxylases by Citric Acid Cycle Intermediates POSSIBLE LINKS BETWEEN CELL METABOLISM AND STABILIZATION OF HIF

    Peppi Koivunen;Maija Hirsilä;Anne M. Remes;Ilmo E. Hassinen

  • Prolyl 4-hydroxylases, the key enzymes of collagen biosynthesis.

    Johanna Myllyharju

  • Catalytic properties of the asparaginyl hydroxylase (FIH) in the oxygen sensing pathway are distinct from those of its prolyl 4-hydroxylases.

    Peppi Koivunen;Maija Hirsilä;Volkmar Günzler;Kari I. Kivirikko

  • Prolyl 4-hydroxylases and their protein disulfide isomerase subunit

    Kari I. Kivirikko;Johanna Myllyharju

  • Lysyl oxidase is essential for normal development and function of the respiratory system and for the integrity of elastic and collagen fibers in various tissues.

    Joni M. Mäki;Raija Sormunen;Sari Lippo;Riitta Kaarteenaho-Wiik

  • Prolyl 4-hydroxylases, key enzymes in the synthesis of collagens and regulation of the response to hypoxia, and their roles as treatment targets

    Johanna Myllyharju

  • Prolyl 4-hydroxylation regulates Argonaute 2 stability

    Hank H. Qi;Pat P. Ongusaha;Johanna Myllyharju;Dongmei Cheng

  • Characterization of the iron‐ and 2‐oxoglutarate‐binding sites of human prolyl 4‐hydroxylase

    Johanna Myllyharju;Kari I. Kivirikko

  • Assembly of human prolyl 4‐hydroxylase and type III collagen in the yeast Pichia pastoris: formation of a stable enzyme tetramer requires coexpression with collagen and assembly of a stable collagen requires coexpression with prolyl 4‐hydroxylase

    Annamari Vuorela;Johanna Myllyharju;Ritva Nissi;Taina Pihlajaniemi

  • Effect of desferrioxamine and metals on the hydroxylases in the oxygen sensing pathway

    Maija Hirsilä;Peppi Koivunen;Leon Xu;Todd Seeley

  • Oxygen-dependent ATF-4 stability is mediated by the PHD3 oxygen sensor

    Jens Köditz;Jutta Nesper;Marieke Wottawa;Daniel P. Stiehl

  • Roles of the human hypoxia-inducible factor (HIF)-3α variants in the hypoxia response

    Minna Heikkilä;Annika Pasanen;Kari I. Kivirikko;Johanna Myllyharju

  • Collagen binding specificity of the discoidin domain receptors: Binding sites on collagens II and III and molecular determinants for collagen IV recognition by DDR1

    Huifang Xu;Nicolas Raynal;Stavros Stathopoulos;Johanna Myllyharju

  • Characterization of Human Type III Collagen Expressed in a Baculovirus System PRODUCTION OF A PROTEIN WITH A STABLE TRIPLE HELIX REQUIRES COEXPRESSION WITH THE TWO TYPES OF RECOMBINANT PROLYL 4-HYDROXYLASE SUBUNIT

    Arja Lamberg;Tarja Helaakoski;Johanna Myllyharju;Sirkku Peltonen

  • Cloning of the Human Prolyl 4-Hydroxylase α Subunit Isoform α(II) and Characterization of the Type II Enzyme Tetramer THE α(I) AND α(II) SUBUNITS DO NOT FORM A MIXED α(I)α(II)β2 TETRAMER

    Pia Annunen;Tarja Helaakoski;Johanna Myllyharju;Johanna Veijola

  • High‐level production of human type I collagen in the yeast Pichia pastoris

    Minna Nokelainen;Hongmin Tu;Annamari Vuorela;Holger Notbohm

  • Cloning and characterization of a low molecular weight prolyl 4-hydroxylase from Arabidopsis thaliana. Effective hydroxylation of proline-rich, collagen-like, and hypoxia-inducible transcription factor alpha-like peptides.

    Reija Hieta;Johanna Myllyharju

Frequent Co-Authors

Kari I. Kivirikko
Kari I. Kivirikko University of Oulu
Raija Sormunen
Raija Sormunen University of Oulu
Peter Neubauer
Peter Neubauer Technical University of Berlin
Taina Pihlajaniemi
Taina Pihlajaniemi University of Oulu
Rik K. Wierenga
Rik K. Wierenga University of Oulu
Peter J. Ratcliffe
Peter J. Ratcliffe The Francis Crick Institute
Attila Aszódi
Attila Aszódi Ludwig-Maximilians-Universität München
Tadhg P. Begley
Tadhg P. Begley Texas A&M University
John M. Stuart
John M. Stuart University of Tennessee Health Science Center
David D. Brand
David D. Brand University of Tennessee Health Science Center

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Best Scientists Citing Johanna Myllyharju

Christopher J. Schofield

Christopher J. Schofield

University of Oxford

Publications: 127

Peter J. Ratcliffe

Peter J. Ratcliffe

The Francis Crick Institute

Publications: 73

Hans Peter Bächinger

Hans Peter Bächinger

Oregon Health & Science University

Publications: 59

William G. Kaelin

William G. Kaelin

Harvard University

Publications: 55

Michael A. McDonough

Michael A. McDonough

University of Oxford

Publications: 37

Barbara Brodsky

Barbara Brodsky

Tufts University

Publications: 35

Taina Pihlajaniemi

Taina Pihlajaniemi

University of Oulu

Publications: 31

Christopher W. Pugh

Christopher W. Pugh

University of Oxford

Publications: 31

Mitsuo Yamauchi

Mitsuo Yamauchi

University of North Carolina at Chapel Hill

Publications: 30

MaryAnn Weis

MaryAnn Weis

University of Washington

Publications: 27

Gregg L. Semenza

Gregg L. Semenza

Johns Hopkins University School of Medicine

Publications: 25

Ronald T. Raines

Ronald T. Raines

MIT

Publications: 24

David R. Eyre

David R. Eyre

University of Washington

Publications: 24

Philip C. Trackman

Philip C. Trackman

Boston University

Publications: 23

Prem Puri

Prem Puri

University College Dublin

Publications: 23

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Leena Bruckner-Tuderman

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