World's Best Scientists 2026 revealed!
Johan Billen

Johan Billen

D-Index & Metrics

Ecology and Evolution

D-Index
44
Citations
8619
World Ranking
5026
National Ranking
62

Johan Billen publication distribution in Ecology and Evolution in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Ecology and Evolution in 2026. The highlighted bar marks where Johan Billen sits on this spectrum.

37–41 publications: 2 scientists 42–46 publications: 9 scientists 47–51 publications: 10 scientists 52–56 publications: 31 scientists 57–61 publications: 56 scientists 62–66 publications: 91 scientists 67–71 publications: 92 scientists 72–76 publications: 127 scientists 77–81 publications: 173 scientists 82–86 publications: 236 scientists 87–91 publications: 227 scientists 92–96 publications: 240 scientists 97–101 publications: 333 scientists 102–106 publications: 280 scientists 107–111 publications: 300 scientists 112–116 publications: 285 scientists 117–121 publications: 305 scientists 122–126 publications: 287 scientists 127–131 publications: 278 scientists 132–136 publications: 289 scientists 137–141 publications: 273 scientists 142–146 publications: 262 scientists 147–151 publications: 246 scientists 152–156 publications: 235 scientists 157–161 publications: 205 scientists 162–166 publications: 199 scientists 167–171 publications: 174 scientists 172–176 publications: 164 scientists 177–181 publications: 173 scientists 182–186 publications: 189 scientists 187–191 publications: 170 scientists 192–196 publications: 139 scientists 197–201 publications: 128 scientists 202–206 publications: 117 scientists 207–211 publications: 115 scientists 212–216 publications: 107 scientists 217–221 publications: 124 scientists 222–226 publications: 81 scientists 227–231 publications: 82 scientists 232–236 publications: 85 scientists 237–241 publications: 75 scientists 242–246 publications: 66 scientists 247–251 publications: 81 scientists 252–256 publications: 69 scientists 257–261 publications: 70 scientists 262–266 publications: 55 scientists 267–271 publications: 64 scientists 272–276 publications: 49 scientists 277–281 publications: 48 scientists 282–286 publications: 44 scientists 287–291 publications: 45 scientists 292–296 publications: 36 scientists 297–301 publications: 39 scientists 302–306 publications: 38 scientists 307–311 publications: 30 scientists 312–316 publications: 34 scientists 317–321 publications: 25 scientists 322–326 publications: 22 scientists 327–331 publications: 28 scientists 332–336 publications: 30 scientists 337–341 publications: 22 scientists 342–346 publications: 30 scientists 347–351 publications: 26 scientists 352–356 publications: 22 scientists 357–361 publications: 29 scientists 362–366 publications: 21 scientists 367–371 publications: 12 scientists 372–376 publications: 18 scientists 377–381 publications: 17 scientists 382–386 publications: 13 scientists 387–391 publications: 21 scientists 392–396 publications: 13 scientists 397–401 publications: 10 scientists 402–406 publications: 15 scientists 407–411 publications: 8 scientists 412–416 publications: 9 scientists 417–421 publications: 8 scientists 422–426 publications: 4 scientists 427–431 publications: 10 scientists 432–436 publications: 10 scientists 437–441 publications: 5 scientists 442–446 publications: 4 scientists 447–451 publications: 10 scientists 452–456 publications: 2 scientists 457–461 publications: 4 scientists 462–466 publications: 7 scientists 467–471 publications: 7 scientists 472–476 publications: 6 scientists 477–481 publications: 6 scientists 482–486 publications: 3 scientists 487–491 publications: 3 scientists 492–496 publications: 4 scientists 497–501 publications: 4 scientists 502–506 publications: 8 scientists 507–511 publications: 5 scientists 512–516 publications: 4 scientists 517–521 publications: 5 scientists 522–526 publications: 4 scientists 527–530 publications: 5 scientists 531+ publications: 100 scientists
37 publications 531+

This scientist: 293 publications — 91st percentile

91% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 531 publications or more.

Johan Billen D-index placement in Ecology and Evolution in 2026

The chart shows the D-index (discipline H-index) distribution of Ecology and Evolution scientists ranked by Research.com in 2026. The highlighted bar marks where Johan Billen sits on this spectrum.

30 D-Index: 95 scientists 31 D-Index: 123 scientists 32 D-Index: 191 scientists 33 D-Index: 245 scientists 34 D-Index: 252 scientists 35 D-Index: 244 scientists 36 D-Index: 249 scientists 37 D-Index: 244 scientists 38 D-Index: 254 scientists 39 D-Index: 254 scientists 40 D-Index: 280 scientists 41 D-Index: 274 scientists 42 D-Index: 266 scientists 43 D-Index: 233 scientists 44 D-Index: 248 scientists 45 D-Index: 208 scientists 46 D-Index: 201 scientists 47 D-Index: 213 scientists 48 D-Index: 207 scientists 49 D-Index: 172 scientists 50 D-Index: 209 scientists 51 D-Index: 199 scientists 52 D-Index: 153 scientists 53 D-Index: 169 scientists 54 D-Index: 163 scientists 55 D-Index: 148 scientists 56 D-Index: 120 scientists 57 D-Index: 146 scientists 58 D-Index: 137 scientists 59 D-Index: 140 scientists 60 D-Index: 104 scientists 61 D-Index: 119 scientists 62 D-Index: 118 scientists 63 D-Index: 99 scientists 64 D-Index: 77 scientists 65 D-Index: 100 scientists 66 D-Index: 86 scientists 67 D-Index: 83 scientists 68 D-Index: 75 scientists 69 D-Index: 94 scientists 70 D-Index: 57 scientists 71 D-Index: 68 scientists 72 D-Index: 59 scientists 73 D-Index: 63 scientists 74 D-Index: 66 scientists 75 D-Index: 60 scientists 76 D-Index: 42 scientists 77 D-Index: 49 scientists 78 D-Index: 39 scientists 79 D-Index: 37 scientists 80 D-Index: 43 scientists 81 D-Index: 41 scientists 82 D-Index: 45 scientists 83 D-Index: 34 scientists 84 D-Index: 40 scientists 85 D-Index: 35 scientists 86 D-Index: 55 scientists 87 D-Index: 24 scientists 88 D-Index: 24 scientists 89 D-Index: 30 scientists 90 D-Index: 23 scientists 91 D-Index: 25 scientists 92 D-Index: 27 scientists 93 D-Index: 27 scientists 94 D-Index: 13 scientists 95 D-Index: 23 scientists 96 D-Index: 12 scientists 97 D-Index: 14 scientists 98 D-Index: 19 scientists 99 D-Index: 16 scientists 100 D-Index: 14 scientists 101 D-Index: 5 scientists 102 D-Index: 21 scientists 103 D-Index: 16 scientists 104 D-Index: 10 scientists 105 D-Index: 7 scientists 106 D-Index: 7 scientists 107 D-Index: 7 scientists 108 D-Index: 6 scientists 109 D-Index: 9 scientists 110 D-Index: 11 scientists 111 D-Index: 12 scientists 112 D-Index: 9 scientists 113 D-Index: 7 scientists 114 D-Index: 3 scientists 115 D-Index: 8 scientists 116 D-Index: 5 scientists 117 D-Index: 11 scientists 118 D-Index: 4 scientists 119 D-Index: 2 scientists 120 D-Index: 5 scientists 121+ D-Index: 100 scientists
30 D-Index 121+

This scientist: 44 D-Index — 41st percentile

41% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 121 D-Index or more.

Overview

Johan Billen is affiliated with KU Leuven in Belgium and has contributed extensively to the fields of Agricultural and Biological Sciences as well as Biochemistry, Genetics and Molecular Biology. Their research encompasses various subfields, including Ecology, Evolution, Behavior and Systematics, Genetics, Insect Science, Cellular and Molecular Neuroscience, and Molecular Biology. The main topics covered in their work include:

  • Insect and Arachnid Ecology and Behavior
  • Plant and animal studies
  • Insect and Pesticide Research
  • Neurobiology and Insect Physiology Research
  • Fossil Insects in Amber
  • Animal Behavior and Reproduction
  • Forest Insect Ecology and Management

Billen has published in numerous scientific venues with varying frequency. The most frequent publication venues are:

  • Arthropod Structure & Development
  • Insectes Sociaux
  • SSRN Electronic Journal
  • Zenodo (CERN European Organization for Nuclear Research)
  • Lirias (KU Leuven)

The scientist's recent papers demonstrate a focus on insect anatomy and molecular biology, including studies on hymenopteran species and RNA interference. Some notable publications include:

  • "Comparative analysis of worker head anatomy of Formica and Brachyponera (Hymenoptera: Formicidae)", 2020, Lirias (KU Leuven)
  • "The head anatomy of Protanilla lini (Hymenoptera: Formicidae: Leptanillinae), with a hypothesis of their mandibular movement", 2021, Lirias (KU Leuven)
  • "Extracellular vesicles spread the RNA interference signal of Tribolium castaneum TcA cells", 2020, Insect Biochemistry and Molecular Biology
  • "Structural diversity of symbionts and related cellular mechanisms underlying vertical symbiont transmission in cicadas", 2021, Environmental Microbiology
  • "The ant abdomen: The skeletomuscular and soft tissue anatomy of Amblyopone australis workers (Hymenoptera: Formicidae)", 2022, Journal of Morphology

Johan Billen frequently collaborates with various researchers. The most regular coauthors include:

  • Chu Wang
  • Chung-Chi Lin
  • Evan P. Economo
  • Adrian Richter
  • Hong He

Best Publications

  • Metformin promotes lifespan through mitohormesis via the peroxiredoxin PRDX-2

    Wouter De Haes;Lotte Frooninckx;Roel Van Assche;Arne Smolders

  • Coevolved Crypts and Exocrine Glands Support Mutualistic Bacteria in Fungus-Growing Ants

    Cameron Robert Currie;Michael Poulsen;John Mendenhall;Jacobus J. Koos Boomsma

  • Conserved class of queen pheromones stops social insect workers from reproducing.

    Annette Van Oystaeyen;Ricardo Caliari Oliveira;Luke Holman;Jelle S. van Zweden

  • The life of a dead ant: the expression of an adaptive extended phenotype.

    Sandra B. Andersen;Sylvia Gerritsma;Kalsum M. Yusah;David Mayntz

  • Behavioral mechanisms and morphological symptoms of zombie ants dying from fungal infection

    David P Hughes;David P Hughes;Sandra B Andersen;Nigel L Hywel-Jones;Winanda Himaman

  • Tetraponera ants have gut symbionts related to nitrogen–fixing root–nodule bacteria

    Steven Van Borm;Alfred Buschinger;Jacobus J. Koos Boomsma;Johan Billen

  • Biology and evolution of social insects

    Johan Billen

  • Widespread occurrence of the micro-organism Wolbachia in ants

    T. Wenseleers;F. Ito;S. Van Borm;R. Huybrechts

  • Age-dependent morphology and ultrastructure of the hypopharyngeal gland of Apis mellifera workers (Hymenoptera, Apidae)

    Jeroen Deseyn;Johan Billen

  • Caste fate conflict in swarm-founding social hymenoptera: an inclusive fitness analysis.

    Tom Wenseleers;Francis L.W. Ratnieks;Johan Billen

  • Wolbachia in Leafcutter Ants: a Widespread Symbiont That May Induce Male Killing or Incompatible Matings

    S. van Borm;T. Wenseleers;J. Billen;Jacobus Jan Boomsma

  • Policing behaviour towards virgin egg layers in a polygynous ponerine ant.

    Bruno Gobin;Johan Billen;Christian Peeters

  • Occurrence and structural organization of the exocrine glands in the legs of ants

    Johan Billen

  • Chemistry of metapleural gland secretions of three attine ants, Atta sexdens rubropilosa, Atta cephalotes , and Acromyrmex octospinosus (Hymenoptera: Formicidae)

    Ruth R. Do Nascimento;Eric Schoeters;E. David Morgan;Johan Billen

  • The diversity of microorganisms associated with Acromyrmex leafcutter ants

    Steven Van Borm;Johan Billen;Jacobus J. Koos Boomsma

  • Deleterious Wolbachia in the ant Formica truncorum

    Tom Wenseleers;Lotta Sundstrom;Johan Billen

  • The importance of the spermathecal duct in bumblebees.

    Eric Schoeters;Johan Billen

  • Task-related variation of postpharyngeal and cuticular hydrocarbon compositions in the ant Myrmicaria eumenoides

    Manfred Kaib;B. Eisermann;E. Schoeters;J. Billen

  • Structural and chemical ontogeny of the postpharyngeal gland in the desert ant Cataglyphis niger

    Victoria Soroker;Abraham Hefetz;M Cojocaru;Johan Billen

  • On the similarity of the Dufour gland secretion and the cuticular hydrocarbons of some bumblebees

    N. J. Oldham;Johan Billen;E. D. Morgan

  • Pheromone Communication in Social Insects: Sources and Secretions

    Johan Billen;E. David Morgan

Frequent Co-Authors

Christian Peeters
Christian Peeters Sorbonne University
Jacobus J. Boomsma
Jacobus J. Boomsma University of Copenhagen
Patrizia d'Ettorre
Patrizia d'Ettorre Paris 13 University
Abraham Hefetz
Abraham Hefetz Tel Aviv University
Falko P. Drijfhout
Falko P. Drijfhout Keele University
Vera Lúcia Imperatriz-Fonseca
Vera Lúcia Imperatriz-Fonseca Universidade de São Paulo
Kazuki Tsuji
Kazuki Tsuji University of the Ryukyus
Jean-Louis Deneubourg
Jean-Louis Deneubourg Université Libre de Bruxelles

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