World's Best Scientists 2026 revealed!

D-Index & Metrics

Biology and Biochemistry

D-Index
57
Citations
13318
World Ranking
13702
National Ranking
5807

Ralph J. Greenspan publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Ralph J. Greenspan sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 139 publications — 21st percentile

21% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Ralph J. Greenspan D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Ralph J. Greenspan sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 57 D-Index — 31st percentile

31% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Research.com Recognitions

  • 1982 - Fellow of Alfred P. Sloan Foundation

Overview

Ralph J. Greenspan is affiliated with the University of California, San Diego in the United States. Their research primarily focuses on neuroscience and related disciplines including biochemistry, genetics, and molecular biology. They have contributed to multiple specialized subfields such as cellular and molecular neuroscience, genetics, ecology, evolution, behavior and systematics, cognitive neuroscience, and endocrine and autonomic systems.

Their scientific work addresses several main topics, notably neurobiology and insect physiology research, insect and arachnid ecology and behavior, plant and animal studies, animal behavior and reproduction, memory and neural mechanisms, circadian rhythm and melatonin, and functional brain connectivity studies.

Ralph J. Greenspan has published research in several reputable journals and venues, including:

  • Nature
  • Journal of Neurogenetics
  • Proceedings of the National Academy of Sciences
  • Nature Communications
  • Frontiers in Behavioral Neuroscience

Some of their recent papers are:

  • Differential mechanisms underlie trace and delay conditioning in Drosophila, 2022, Nature
  • Valence opponency in peripheral olfactory processing, 2022, Proceedings of the National Academy of Sciences
  • Imaging brain activity during complex social behaviors in Drosophila with Flyception2, 2020, Nature Communications
  • Anti-instinctive Learning Behavior Revealed by Locomotion-Triggered Mild Heat Stress in Drosophila, 2020, Frontiers in Behavioral Neuroscience
  • Investigating the impact of electroconvulsive therapy on brain networks and sleep: an observational study protocol, 2025, BMJ Open

They have collaborated frequently with several researchers, including:

  • Dhruv Grover
  • Jen-Yung Chen
  • Jinfang Li
  • Jiayun Xie
  • Jean-Pierre Changeux

Ralph J. Greenspan was recognized as a Fellow of the Alfred P. Sloan Foundation in 1982.

Best Publications

  • Courtship in Drosophila.

    Ralph J. Greenspan;Jean-François Ferveur

  • Natural behavior polymorphism due to a cGMP-dependent protein kinase of Drosophila.

    K. A. Osborne;A. Robichon;A. Robichon;E. Burgess;E. Burgess;S. Butland;S. Butland

  • Stress response genes protect against lethal effects of sleep deprivation in Drosophila.

    Paul J. Shaw;Giulio Tononi;Giulio Tononi;Ralph J. Greenspan;Donald F. Robinson

  • Dopaminergic Modulation of Arousal in Drosophila

    Rozi Andretic;Bruno van Swinderen;Ralph J. Greenspan

  • Fly pushing : the theory and practice of Drosophila genetics

    Ralph J. Greenspan

  • Serotonin and neuropeptide F have opposite modulatory effects on fly aggression.

    Herman A Dierick;Ralph J Greenspan

  • Electrophysiological correlates of rest and activity in Drosophila melanogaster.

    Douglas A Nitz;Bruno van Swinderen;Giulio Tononi;Ralph J Greenspan

  • A Proposal for a Coordinated Effort for the Determination of Brainwide Neuroanatomical Connectivity in Model Organisms at a Mesoscopic Scale

    Jason W. Bohland;Caizhi Wu;Helen Barbas;Hemant Bokil

  • Expression pattern of Motch, a mouse homolog of Drosophila Notch, suggests an important role in early postimplantation mouse development.

    F.F. Del Amo;D.E. Smith;P.J. Swiatek;M. Gendron-Maguire

  • Inhibition of calcium/calmodulin-dependent protein kinase in drosophila disrupts behavioral plasticity

    Leslie C. Griffith;Lynne M. Verselis;Kay Marie Aitken;Charalambos P. Kyriacou

  • Activation of EGFR and ERK by rhomboid signaling regulates the consolidation and maintenance of sleep in Drosophila.

    Krisztina Foltenyi;Ralph J Greenspan;John W Newport

  • Genetic feminization of brain structures and changed sexual orientation in male Drosophila

    Jean-François Ferveur;Klemens F. Störtkuhl;Reinhard F. Stocker;Ralph J. Greenspan

  • Molecular analysis of flies selected for aggressive behavior.

    Herman A Dierick;Ralph J Greenspan

  • Polyspecificity of T cell and B cell receptor recognition

    Kai W. Wucherpfennig;Paul M. Allen;Franco Celada;Irun R. Cohen

  • Genetic feminization of pheromones and its behavioral consequences in Drosophila males.

    Jean-François Ferveur;Fabrice Savarit;Fabrice Savarit;Fabrice Savarit;Cahir J. O'Kane;Cahir J. O'Kane;Cahir J. O'Kane;Gilles Sureau;Gilles Sureau;Gilles Sureau

  • The flexible genome.

    Ralph J. Greenspan

  • Acetylcholinesterase mutants in drosophila and their effects on the structure and function of the cental nervous system

    Ralph J. Greenspan;James A. Finn;Jeffrey C. Hall

  • Identification of genes involved in Drosophila melanogaster geotaxis, a complex behavioral trait

    Daniel P. Toma;Kevin P. White;Jerry Hirsch;Ralph J. Greenspan

  • A Comprehensive Wiring Diagram of the Protocerebral Bridge for Visual Information Processing in the Drosophila Brain

    Chih-Yung Lin;Chao-Chun Chuang;Tzu-En Hua;Chun-Chao Chen

  • Natural Behavior Polymorphism Due to a cGMP-Dependent Protein Kinase of

    Osborne Ka;A. Robichon;Shaw Ra;H. S. Pereira

Frequent Co-Authors

Giulio Tononi
Giulio Tononi University of Wisconsin–Madison
Paul J. Shaw
Paul J. Shaw Washington University in St. Louis
Terrence J. Sejnowski
Terrence J. Sejnowski Salk Institute for Biological Studies
Rafael Yuste
Rafael Yuste Columbia University
Karl Deisseroth
Karl Deisseroth Stanford University
Michael L. Roukes
Michael L. Roukes California Institute of Technology
Bruno van Swinderen
Bruno van Swinderen University of Queensland
George M. Church
George M. Church Harvard University
A. Paul Alivisatos
A. Paul Alivisatos University of Chicago
Leslie C. Griffith
Leslie C. Griffith Brandeis University

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