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Francis S. Willard

D-Index & Metrics

Biology and Biochemistry

D-Index
50
Citations
8749
World Ranking
17661
National Ranking
7246

Francis S. Willard publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Francis S. Willard sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 116 publications — 10th percentile

10% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Francis S. Willard D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Francis S. Willard sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 50 D-Index — 11th percentile

11% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Overview

Francis S. Willard is affiliated with Eli Lilly in the United States and has contributed extensively to research in the fields of medicine and biochemistry, genetics, and molecular biology. Their work predominantly focuses on molecular biology, endocrinology, diabetes and metabolism, surgery, cellular and molecular neuroscience, and pharmacology.

The scientist's research topics include diabetes treatment and management, receptor mechanisms and signaling, pancreatic function and diabetes, neuropeptides and animal physiology, protein kinase regulation and GTPase signaling, pharmacology and obesity treatment, and metabolism related to diabetes and cancer.

Recent publications authored or involving Francis S. Willard include:

  • "Tirzepatide is an imbalanced and biased dual GIP and GLP-1 receptor agonist" (2020, JCI Insight)
  • "LY3437943, a novel triple glucagon, GIP, and GLP-1 receptor agonist for glycemic control and weight loss: From discovery to clinical proof of concept" (2022, Cell Metabolism)
  • "Activation of the GLP-1 receptor by a non-peptidic agonist" (2020, Nature)
  • "Structural basis for GLP-1 receptor activation by LY3502970, an orally active nonpeptide agonist" (2020, Proceedings of the National Academy of Sciences)
  • "The incretin co-agonist tirzepatide requires GIPR for hormone secretion from human islets" (2023, Nature Metabolism)

Frequent co-authors working alongside Willard include:

  • David P. Siderovski
  • Kyle W. Sloop
  • Adam J. Kimple
  • Aaron D. Showalter
  • David B. Wainscott

The most common venues for Willard's publications are:

  • UNC Libraries
  • Diabetes
  • Proceedings of the National Academy of Sciences
  • Biochemical and Biophysical Research Communications
  • JCI Insight

Willard's research integrates detailed studies of receptor mechanisms, particularly involving glucagon-like peptide-1 (GLP-1) and glucose-dependent insulinotropic polypeptide (GIP) receptors, which play essential roles in glycemic control and metabolic regulation.

Best Publications

  • G-protein signaling: back to the future

    C. R. McCudden;M. D. Hains;R. J. Kimple;D. P. Siderovski

  • The GAPs, GEFs, and GDIs of heterotrimeric G-protein alpha subunits.

    David P. Siderovski;Francis S. Willard

  • A Seven-Transmembrane RGS Protein That Modulates Plant Cell Proliferation

    Jin Gui Chen;Francis S. Willard;Jirong Huang;Jiansheng Liang

  • Translation of Polarity Cues into Asymmetric Spindle Positioning in Caenorhabditis elegans Embryos.

    Kelly Colombo;Stephan W. Grill;Randall J. Kimple;Francis S. Willard

  • Crystal structure of the human PRMT5:MEP50 complex

    Stephen Antonysamy;Zahid Bonday;Robert M. Campbell;Brandon Doyle

  • Tirzepatide is an imbalanced and biased dual GIP and GLP-1 receptor agonist

    Francis S. Willard;Jonathan D. Douros;Maria B.N. Gabe;Aaron D. Showalter

  • GTPase acceleration as the rate-limiting step in Arabidopsis G protein-coupled sugar signaling

    Christopher A. Johnston;J. Philip Taylor;Yajun Gao;Adam J. Kimple

  • Return of the GDI: The GoLoco Motif in Cell Division

    Francis S. Willard;Randall J. Kimple;David P. Siderovski

  • Structural diversity in the RGS domain and its interaction with heterotrimeric G protein alpha-subunits.

    Meera Soundararajan;Francis S. Willard;Adam J. Kimple;Andrew P. Turnbull

  • RIC-8 Is Required for GPR-1/2-Dependent Gα Function during Asymmetric Division of C. elegans Embryos

    Katayoun Afshar;Francis S. Willard;Kelly Colombo;Christopher A. Johnston

  • Rgs1 regulates multiple Gα subunits in Magnaporthe pathogenesis, asexual growth and thigmotropism

    Hao Liu;Angayarkanni Suresh;Francis S Willard;David P Siderovski

  • Role of the p70S6K pathway in regulating the actin cytoskeleton and cell migration

    Leise A Berven;Francis S Willard;Michael F Crouch

  • Novel Small Molecule Glucagon-Like Peptide-1 Receptor Agonist Stimulates Insulin Secretion in Rodents and From Human Islets

    Kyle W. Sloop;Francis S. Willard;Martin B. Brenner;James Ficorilli

  • Comment on "A G Protein–Coupled Receptor Is a Plasma Membrane Receptor for the Plant Hormone Abscisic Acid"

    Christopher A. Johnston;Brenda R. Temple;Jin Gui Chen;Yajun Gao

  • Activation of the GLP-1 receptor by a non-peptidic agonist

    Peishen Zhao;Yi Lynn Liang;Matthew J. Belousoff;Giuseppe Deganutti

  • Chronic olanzapine treatment causes differential expression of genes in frontal cortex of rats as revealed by DNA microarray technique.

    S Hossein Fatemi;Teri J Reutiman;Timothy D Folsom;Christopher Bell

  • Structural basis for GLP-1 receptor activation by LY3502970, an orally active nonpeptide agonist.

    Takahiro Kawai;Bingfa Sun;Hitoshi Yoshino;Dan Feng

  • MEK, ERK, and p90RSK are present on mitotic tubulin in Swiss 3T3 cells: A role for the MAP kinase pathway in regulating mitotic exit

    Francis S. Willard;Michael F. Crouch

  • Regulators of G-protein Signaling accelerate GPCR signaling kinetics and govern sensitivity solely by accelerating GTPase activity

    Nevin A. Lambert;Christopher A. Johnston;Christopher A. Johnston;Steven D. Cappell;Sudhakiranmayi Kuravi

  • Cortical localization of the Gα protein GPA-16 requires RIC-8 function during C. elegans asymmetric cell division

    Katayoun Afshar;Francis S. Willard;Kelly Colombo;David P. Siderovski

Frequent Co-Authors

David P. Siderovski
David P. Siderovski West Virginia University
Denise Wootten
Denise Wootten Monash University
Patrick M. Sexton
Patrick M. Sexton Monash University
Arthur Christopoulos
Arthur Christopoulos Monash University
Pierre Gönczy
Pierre Gönczy École Polytechnique Fédérale de Lausanne
Anton Simeonov
Anton Simeonov National Institutes of Health
Ajit Jadhav
Ajit Jadhav Pfizer (Germany)
James Inglese
James Inglese National Institutes of Health
Mischa Machius
Mischa Machius The University of Texas Southwestern Medical Center
Alan M. Jones
Alan M. Jones University of North Carolina at Chapel Hill

External Links

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Best Scientists Citing Francis S. Willard

Alan M. Jones

Alan M. Jones

University of North Carolina at Chapel Hill

Publications: 71

Patrick M. Sexton

Patrick M. Sexton

Monash University

Publications: 50

David P. Siderovski

David P. Siderovski

West Virginia University

Publications: 49

Denise Wootten

Denise Wootten

Monash University

Publications: 45

Sona Pandey

Sona Pandey

Donald Danforth Plant Science Center

Publications: 32

Arthur Christopoulos

Arthur Christopoulos

Monash University

Publications: 31

Ming-Wei Wang

Ming-Wei Wang

Chinese Academy of Sciences

Publications: 31

Sarah M. Assmann

Sarah M. Assmann

Pennsylvania State University

Publications: 29

Jin-Gui Chen

Jin-Gui Chen

Oak Ridge National Laboratory

Publications: 27

John R. Hepler

John R. Hepler

Emory University

Publications: 25

Richard R. Neubig

Richard R. Neubig

Michigan State University

Publications: 24

Stephen M. Lanier

Stephen M. Lanier

Wayne State University

Publications: 23

José Ramón Botella

José Ramón Botella

University of Queensland

Publications: 22

Laurence J. Miller

Laurence J. Miller

Mayo Clinic

Publications: 21

Peter J. Little

Peter J. Little

University of the Sunshine Coast

Publications: 19

Kirill A. Martemyanov

Kirill A. Martemyanov

Scripps Research Institute

Publications: 17

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