World's Best Scientists 2026 revealed!

D-Index & Metrics

Discipline name D-Index World Ranking National Ranking Publications Citations
Ecology and Evolution 100 279 113 534 30858
Genetics 98 818 409 432 28164

Francisco J. Ayala publications per year

The chart shows the history of publications by Francisco J. Ayala between 1966 and 2025, highlighting the no. of papers published in each year and offering an overview of the publication velocity of this scholar. Francisco J. Ayala published across 60 years, from 1966 to 2025, averaging 10.7 papers a year. Output peaked at 38 publications in 2008. 3 of the 639 publications appeared in the last two years.

No. of publications
10 20 30
Bar chart. Horizontal axis: year, 1966 to 2025. Vertical axis: number of publications, 0 to 38. Peak 38 publications in 2008. 1966: 3 publications 1967: 2 publications 1968: 2 publications 1969: 4 publications 1970: 3 publications 1971: 2 publications 1972: 8 publications 1973: 8 publications 1974: 16 publications 1975: 12 publications 1976: 8 publications 1977: 7 publications 1978: 9 publications 1979: 7 publications 1980: 4 publications 1981: 11 publications 1982: 9 publications 1983: 5 publications 1984: 1 publication 1985: 6 publications 1986: 13 publications 1987: 6 publications 1988: 7 publications 1989: 14 publications 1990: 4 publications 1991: 11 publications 1992: 6 publications 1993: 5 publications 1994: 12 publications 1995: 16 publications 1996: 13 publications 1997: 10 publications 1998: 10 publications 1999: 15 publications 2000: 12 publications 2001: 8 publications 2002: 9 publications 2003: 13 publications 2004: 10 publications 2005: 12 publications 2006: 14 publications 2007: 23 publications 2008: 38 publications 2009: 37 publications 2010: 31 publications 2011: 16 publications 2012: 10 publications 2013: 19 publications 2014: 20 publications 2015: 15 publications 2016: 24 publications 2017: 29 publications 2018: 9 publications 2019: 4 publications 2020: 3 publications 2021: 4 publications 2022: 3 publications 2023: 4 publications 2024: 1 publication 2025: 2 publications
1966 2025

639 publications in total across all disciplines

View publications per year as a table
Francisco J. Ayala: publications per year, 1966 to 2025
Year Publications
1966 3
1967 2
1968 2
1969 4
1970 3
1971 2
1972 8
1973 8
1974 16
1975 12
1976 8
1977 7
1978 9
1979 7
1980 4
1981 11
1982 9
1983 5
1984 1
1985 6
1986 13
1987 6
1988 7
1989 14
1990 4
1991 11
1992 6
1993 5
1994 12
1995 16
1996 13
1997 10
1998 10
1999 15
2000 12
2001 8
2002 9
2003 13
2004 10
2005 12
2006 14
2007 23
2008 38
2009 37
2010 31
2011 16
2012 10
2013 19
2014 20
2015 15
2016 24
2017 29
2018 9
2019 4
2020 3
2021 4
2022 3
2023 4
2024 1
2025 2
Total 639
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Francisco J. Ayala publication distribution in Ecology and Evolution in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Ecology and Evolution in 2026. The highlighted bar marks where Francisco J. Ayala sits on this spectrum.

No. of scientists
100 200 300
Bar chart with 100 bars. Horizontal axis: publications, 37–41 to 531+. Vertical axis: number of scientists, 0 to 333. Most scientists, 333, have 97–101 publications. The last bar groups every scientist with 531 publications or more. The highlighted bar, 531+ publications, is where this scientist sits. 37–41 publications: 2 scientists 42–46 publications: 9 scientists 47–51 publications: 10 scientists 52–56 publications: 31 scientists 57–61 publications: 56 scientists 62–66 publications: 91 scientists 67–71 publications: 92 scientists 72–76 publications: 127 scientists 77–81 publications: 173 scientists 82–86 publications: 236 scientists 87–91 publications: 227 scientists 92–96 publications: 240 scientists 97–101 publications: 333 scientists 102–106 publications: 280 scientists 107–111 publications: 300 scientists 112–116 publications: 285 scientists 117–121 publications: 305 scientists 122–126 publications: 287 scientists 127–131 publications: 278 scientists 132–136 publications: 289 scientists 137–141 publications: 273 scientists 142–146 publications: 262 scientists 147–151 publications: 246 scientists 152–156 publications: 235 scientists 157–161 publications: 205 scientists 162–166 publications: 199 scientists 167–171 publications: 174 scientists 172–176 publications: 164 scientists 177–181 publications: 173 scientists 182–186 publications: 189 scientists 187–191 publications: 170 scientists 192–196 publications: 139 scientists 197–201 publications: 128 scientists 202–206 publications: 117 scientists 207–211 publications: 115 scientists 212–216 publications: 107 scientists 217–221 publications: 124 scientists 222–226 publications: 81 scientists 227–231 publications: 82 scientists 232–236 publications: 85 scientists 237–241 publications: 75 scientists 242–246 publications: 66 scientists 247–251 publications: 81 scientists 252–256 publications: 69 scientists 257–261 publications: 70 scientists 262–266 publications: 55 scientists 267–271 publications: 64 scientists 272–276 publications: 49 scientists 277–281 publications: 48 scientists 282–286 publications: 44 scientists 287–291 publications: 45 scientists 292–296 publications: 36 scientists 297–301 publications: 39 scientists 302–306 publications: 38 scientists 307–311 publications: 30 scientists 312–316 publications: 34 scientists 317–321 publications: 25 scientists 322–326 publications: 22 scientists 327–331 publications: 28 scientists 332–336 publications: 30 scientists 337–341 publications: 22 scientists 342–346 publications: 30 scientists 347–351 publications: 26 scientists 352–356 publications: 22 scientists 357–361 publications: 29 scientists 362–366 publications: 21 scientists 367–371 publications: 12 scientists 372–376 publications: 18 scientists 377–381 publications: 17 scientists 382–386 publications: 13 scientists 387–391 publications: 21 scientists 392–396 publications: 13 scientists 397–401 publications: 10 scientists 402–406 publications: 15 scientists 407–411 publications: 8 scientists 412–416 publications: 9 scientists 417–421 publications: 8 scientists 422–426 publications: 4 scientists 427–431 publications: 10 scientists 432–436 publications: 10 scientists 437–441 publications: 5 scientists 442–446 publications: 4 scientists 447–451 publications: 10 scientists 452–456 publications: 2 scientists 457–461 publications: 4 scientists 462–466 publications: 7 scientists 467–471 publications: 7 scientists 472–476 publications: 6 scientists 477–481 publications: 6 scientists 482–486 publications: 3 scientists 487–491 publications: 3 scientists 492–496 publications: 4 scientists 497–501 publications: 4 scientists 502–506 publications: 8 scientists 507–511 publications: 5 scientists 512–516 publications: 4 scientists 517–521 publications: 5 scientists 522–526 publications: 4 scientists 527–530 publications: 5 scientists 531+ publications: 100 scientists
37–41 publications 531+

This scientist: 534 publications — 99th percentile

99% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 531 publications or more.

View publications distribution as a table
Number of Ecology and Evolution scientists by publication count, Research.com 2026 ranking edition. Based on 8,381 ranked scientists.
Publications Scientists This scientist
37–41 2
42–46 9
47–51 10
52–56 31
57–61 56
62–66 91
67–71 92
72–76 127
77–81 173
82–86 236
87–91 227
92–96 240
97–101 333
102–106 280
107–111 300
112–116 285
117–121 305
122–126 287
127–131 278
132–136 289
137–141 273
142–146 262
147–151 246
152–156 235
157–161 205
162–166 199
167–171 174
172–176 164
177–181 173
182–186 189
187–191 170
192–196 139
197–201 128
202–206 117
207–211 115
212–216 107
217–221 124
222–226 81
227–231 82
232–236 85
237–241 75
242–246 66
247–251 81
252–256 69
257–261 70
262–266 55
267–271 64
272–276 49
277–281 48
282–286 44
287–291 45
292–296 36
297–301 39
302–306 38
307–311 30
312–316 34
317–321 25
322–326 22
327–331 28
332–336 30
337–341 22
342–346 30
347–351 26
352–356 22
357–361 29
362–366 21
367–371 12
372–376 18
377–381 17
382–386 13
387–391 21
392–396 13
397–401 10
402–406 15
407–411 8
412–416 9
417–421 8
422–426 4
427–431 10
432–436 10
437–441 5
442–446 4
447–451 10
452–456 2
457–461 4
462–466 7
467–471 7
472–476 6
477–481 6
482–486 3
487–491 3
492–496 4
497–501 4
502–506 8
507–511 5
512–516 4
517–521 5
522–526 4
527–530 5
531+ 100 534
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Francisco J. Ayala D-index placement in Ecology and Evolution in 2026

The chart shows the D-index (discipline H-index) distribution of Ecology and Evolution scientists ranked by Research.com in 2026. The highlighted bar marks where Francisco J. Ayala sits on this spectrum.

No. of scientists
50 100 150 200 250
Bar chart with 92 bars. Horizontal axis: D-Index, 30 to 121+. Vertical axis: number of scientists, 0 to 280. Most scientists, 280, have 40 D-Index. The last bar groups every scientist with 121 D-Index or more. The highlighted bar, 100 D-Index, is where this scientist sits. 30 D-Index: 95 scientists 31 D-Index: 123 scientists 32 D-Index: 191 scientists 33 D-Index: 245 scientists 34 D-Index: 252 scientists 35 D-Index: 244 scientists 36 D-Index: 249 scientists 37 D-Index: 244 scientists 38 D-Index: 254 scientists 39 D-Index: 254 scientists 40 D-Index: 280 scientists 41 D-Index: 274 scientists 42 D-Index: 266 scientists 43 D-Index: 233 scientists 44 D-Index: 248 scientists 45 D-Index: 208 scientists 46 D-Index: 201 scientists 47 D-Index: 213 scientists 48 D-Index: 207 scientists 49 D-Index: 172 scientists 50 D-Index: 209 scientists 51 D-Index: 199 scientists 52 D-Index: 153 scientists 53 D-Index: 169 scientists 54 D-Index: 163 scientists 55 D-Index: 148 scientists 56 D-Index: 120 scientists 57 D-Index: 146 scientists 58 D-Index: 137 scientists 59 D-Index: 140 scientists 60 D-Index: 104 scientists 61 D-Index: 119 scientists 62 D-Index: 118 scientists 63 D-Index: 99 scientists 64 D-Index: 77 scientists 65 D-Index: 100 scientists 66 D-Index: 86 scientists 67 D-Index: 83 scientists 68 D-Index: 75 scientists 69 D-Index: 94 scientists 70 D-Index: 57 scientists 71 D-Index: 68 scientists 72 D-Index: 59 scientists 73 D-Index: 63 scientists 74 D-Index: 66 scientists 75 D-Index: 60 scientists 76 D-Index: 42 scientists 77 D-Index: 49 scientists 78 D-Index: 39 scientists 79 D-Index: 37 scientists 80 D-Index: 43 scientists 81 D-Index: 41 scientists 82 D-Index: 45 scientists 83 D-Index: 34 scientists 84 D-Index: 40 scientists 85 D-Index: 35 scientists 86 D-Index: 55 scientists 87 D-Index: 24 scientists 88 D-Index: 24 scientists 89 D-Index: 30 scientists 90 D-Index: 23 scientists 91 D-Index: 25 scientists 92 D-Index: 27 scientists 93 D-Index: 27 scientists 94 D-Index: 13 scientists 95 D-Index: 23 scientists 96 D-Index: 12 scientists 97 D-Index: 14 scientists 98 D-Index: 19 scientists 99 D-Index: 16 scientists 100 D-Index: 14 scientists 101 D-Index: 5 scientists 102 D-Index: 21 scientists 103 D-Index: 16 scientists 104 D-Index: 10 scientists 105 D-Index: 7 scientists 106 D-Index: 7 scientists 107 D-Index: 7 scientists 108 D-Index: 6 scientists 109 D-Index: 9 scientists 110 D-Index: 11 scientists 111 D-Index: 12 scientists 112 D-Index: 9 scientists 113 D-Index: 7 scientists 114 D-Index: 3 scientists 115 D-Index: 8 scientists 116 D-Index: 5 scientists 117 D-Index: 11 scientists 118 D-Index: 4 scientists 119 D-Index: 2 scientists 120 D-Index: 5 scientists 121+ D-Index: 100 scientists
30 D-Index 121+

This scientist: 100 D-Index — 97th percentile

97% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 121 D-Index or more.

View D-Index distribution as a table
Number of Ecology and Evolution scientists by D-index, Research.com 2026 ranking edition. Based on 8,381 ranked scientists.
D-Index Scientists This scientist
30 95
31 123
32 191
33 245
34 252
35 244
36 249
37 244
38 254
39 254
40 280
41 274
42 266
43 233
44 248
45 208
46 201
47 213
48 207
49 172
50 209
51 199
52 153
53 169
54 163
55 148
56 120
57 146
58 137
59 140
60 104
61 119
62 118
63 99
64 77
65 100
66 86
67 83
68 75
69 94
70 57
71 68
72 59
73 63
74 66
75 60
76 42
77 49
78 39
79 37
80 43
81 41
82 45
83 34
84 40
85 35
86 55
87 24
88 24
89 30
90 23
91 25
92 27
93 27
94 13
95 23
96 12
97 14
98 19
99 16
100 14 100
101 5
102 21
103 16
104 10
105 7
106 7
107 7
108 6
109 9
110 11
111 12
112 9
113 7
114 3
115 8
116 5
117 11
118 4
119 2
120 5
121+ 100
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Research.com Recognitions

  • 2017 - Member of the European Academy of Sciences
  • 2001 - US President's National Medal of Science "For his theoretical and experimental discoveries on the origin of species, genetic diversity, and population dynamics that led to a new understanding of biological evolution, and his distinguished contributions to education, the promotion of public understanding of science, and the philosophy and ethics of the scientific enterprise.", Presented by President George W. Bush in a White House East Room ceremony on June 12, 2002.
  • 1980 - Member of the National Academy of Sciences
  • 1977 - Fellow of John Simon Guggenheim Memorial Foundation
  • 1977 - Fellow of the American Academy of Arts and Sciences
  • 1968 - Fellow of the American Association for the Advancement of Science (AAAS)

Overview

Francisco J. Ayala was affiliated with the University of California, Irvine in the United States. Their research focused primarily on fields related to Medicine and Biochemistry, Genetics and Molecular Biology, with significant contributions specifically in Oncology, Cancer Research, Pulmonary and Respiratory Medicine, Cardiology and Cardiovascular Medicine, and Genetics.

They authored numerous papers covering topics that included Breast Cancer Treatment Studies, Advanced Breast Cancer Therapies, Cancer Immunotherapy and Biomarkers, HER2/EGFR in Cancer Research, Cancer Treatment and Pharmacology, Inflammatory Biomarkers in Disease Prognosis, and the Economic and Financial Impacts of Cancer.

Their recent publications included:

  • Mitochondrial DNA drives abscopal responses to radiation that are inhibited by autophagy, 2020, Nature Immunology
  • Mechanobiology of Autophagy: The Unexplored Side of Cancer, 2021, Frontiers in Oncology
  • SEOM-GEICAM-SOLTI clinical guidelines for early-stage breast cancer (2022), 2023, Clinical & Translational Oncology
  • Estratificación, monitorización y control del riesgo cardiovascular en pacientes con cáncer. Documento de consenso de SEC, FEC, SEOM, SEOR, SEHH, SEMG, AEEMT, AEEC y AECC, 2021, Revista Española de Cardiología
  • Assessment of a Genomic Assay in Patients With ERBB2-Positive Breast Cancer Following Neoadjuvant Trastuzumab-Based Chemotherapy With or Without Pertuzumab, 2023, JAMA Oncology

Frequent co-authors included:

  • Esmeralda García-Torralba
  • Esther Navarro Manzano
  • Pilar de la Morena Barrio
  • Sara López-Tarruella
  • Alejandra Ivars Rubio

Their work was frequently published in venues such as Annals of Oncology, Clinical & Translational Oncology, Cancer Research, Journal of Clinical Oncology, and Clinical Cancer Research.

Throughout their career, Francisco J. Ayala received several acknowledgments, including membership in the European Academy of Sciences in 2017 and the National Academy of Sciences in 1980. They were also a Fellow of the John Simon Guggenheim Memorial Foundation and the American Academy of Arts and Sciences, both from 1977, as well as a Fellow of the American Association for the Advancement of Science (AAAS) since 1968.

In 2001, they were awarded the US President's National Medal of Science for theoretical and experimental discoveries on the origin of species, genetic diversity, and population dynamics, along with their contributions to education, public understanding of science, and the philosophy and ethics of the scientific enterprise. This award was presented by President George W. Bush in 2002.

Best Publications

  • ENZYME VARIABILITY IN THE DROSOPHILA WILLISTONI GROUP. IV. GENIC VARIATION IN NATURAL POPULATIONS OF DROSOPHILA WILLISTONI

    Francisco J. Ayala;Jeffrey R. Powell;Martin L. Tracey;Celso A. Mourão

  • Global Models of Growth and Competition

    Michael E. Gilpin;Francisco J. Ayala

  • A clonal theory of parasitic protozoa: the population structures of Entamoeba, Giardia, Leishmania, Naegleria, Plasmodium, Trichomonas, and Trypanosoma and their medical and taxonomical consequences

    Michel Tibayrenc;Finn Kjellberg;Francisco J. Ayala

  • Pseudogenes: are they "junk" or functional DNA?

    Evgeniy S. Balakirev;Francisco J. Ayala

  • Evolutionary and geographical history of the Leishmania donovani complex with a revision of current taxonomy

    Julius Lukeš;Isabel L. Mauricio;Gabriele Schönian;Jean Claude Dujardin

  • Teleological Explanations in Evolutionary Biology

    Francisco J. Ayala

  • Studies in the Philosophy of Biology

    Francisco Jose Ayala;Theodosius Dobzhansky

  • Natural populations of Trypanosoma cruzi, the agent of Chagas disease, have a complex multiclonal structure

    Michel Tibayrenc;Philip Ward;Andres Moya;Francisco J. Ayala

  • Frequency-Dependent Selection

    Francisco J. Ayala;Cathryn A. Campbell

  • Chromosome speciation: Humans, Drosophila, and mosquitoes

    Francisco J. Ayala;Mario Coluzzi

  • Origin of the metazoan phyla: Molecular clocks confirm paleontological estimates

    Ayala Fj;Rzhetsky A

  • Are eukaryotic microorganisms clonal or sexual? A population genetics vantage.

    M Tibayrenc;F Kjellberg;J Arnaud;B Oury

  • Competition between species: theoretical models and experimental tests.

    Francisco J. Ayala;Michael E. Gilpin;Joan G. Ehrenfeld

  • Genetic characterization of six parasitic protozoa: parity between random-primer DNA typing and multilocus enzyme electrophoresis.

    Michel Tibayrenc;Katja Neubauer;Christian Barnabe;Francoise Guerrini

  • Phylogeny of the malarial genus Plasmodium, derived from rRNA gene sequences

    Ananias A. Escalante;Francisco J. Ayala

  • Nucleotide sequences provide evidence of genetic exchange among distantly related lineages of Trypanosoma cruzi

    Carlos A. Machado;Francisco J. Ayala

  • GENETIC DIFFERENTIATION DURING THE SPECIATION PROCESS IN DROSOPHILA.

    Francisco J. Ayala;Martin L. Tracey;Dennis Hedgecock;Rollin C. Richmond

  • Allozymes as Diagnostic Characters of Sibling Species of Drosophila

    Francisco J. Ayala;Jeffrey R. Powell

  • ISOZYME VARIABILITY IN TRYPANOSOMA CRUZI, THE AGENT OF CHAGAS' DISEASE: GENETICAL, TAXONOMICAL, AND EPIDEMIOLOGICAL SIGNIFICANCE.

    Michel Tibayrenc;Francisco J. Ayala

  • Evidence for positive selection in the superoxide dismutase (Sod) region of Drosophila melanogaster.

    Richard R. Hudson;Kevin Bailey;Douglas Skarecky;Jan Kwiatowski

  • Tempo and mode in evolution.

    W M Fitch;F J Ayala

  • Dynamics of Origination and Extinction in the Marine Fossil Record

    John C. Avise;Stephen P. Hubbell;Francisco J. Ayala

  • Homage to Linnaeus: How Many Parasites? How Many Hosts?

    John C. Avise;Stephen P. Hubbell;Francisco J. Ayala

  • Microbes on Mountainsides: Contrasting Elevational Patterns of Bacterial and Plant Diversity

    John C. Avise;Stephen P. Hubbell;Francisco J. Ayala

  • Ecological Extinction and Evolution in the Brave New Ocean

    John C. Avise;Stephen P. Hubbell;Francisco J. Ayala

Frequent Co-Authors

John C. Avise
John C. Avise University of California, Irvine
Michel Tibayrenc
Michel Tibayrenc Institut de Recherche pour le Développement
Stephen P. Hubbell
Stephen P. Hubbell University of California, Los Angeles
François Renaud
François Renaud University of Montpellier
James W. Valentine
James W. Valentine University of California, Berkeley
Andrés Moya
Andrés Moya University of Valencia
Walter M. Fitch
Walter M. Fitch University of California, Irvine
Laurence D. Mueller
Laurence D. Mueller University of California, Irvine
John F. McDonald
John F. McDonald Georgia Institute of Technology
Julius Lukeš
Julius Lukeš Czech Academy of Sciences

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