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Ido Pen

D-Index & Metrics

Discipline name D-Index World Ranking Current World Ranking National Ranking Current National Ranking Publications Citations
Ecology and Evolution 43 5313 5130 142 140 108 7212

Ido Pen publications per year

The chart shows the history of publications by Ido Pen between 1995 and 2025, highlighting the no. of papers published in each year and offering an overview of the publication velocity of this scholar. Ido Pen published across 31 years, from 1995 to 2025, averaging 4.4 papers a year. Output peaked at 11 publications in 2021. 12 of the 135 publications appeared in the last two years.

No. of publications
2 4 6 8 10
Bar chart. Horizontal axis: year, 1995 to 2025. Vertical axis: number of publications, 0 to 11. Peak 11 publications in 2021. 1995: 1 publication 1996: 1 publication 1997: 0 publications 1998: 1 publication 1999: 4 publications 2000: 8 publications 2001: 3 publications 2002: 9 publications 2003: 1 publication 2004: 1 publication 2005: 5 publications 2006: 5 publications 2007: 9 publications 2008: 4 publications 2009: 3 publications 2010: 9 publications 2011: 7 publications 2012: 9 publications 2013: 3 publications 2014: 1 publication 2015: 2 publications 2016: 3 publications 2017: 4 publications 2018: 1 publication 2019: 1 publication 2020: 3 publications 2021: 11 publications 2022: 9 publications 2023: 5 publications 2024: 9 publications 2025: 3 publications
1995 2025

135 publications in total across all disciplines

View publications per year as a table
Ido Pen: publications per year, 1995 to 2025
Year Publications
1995 1
1996 1
1997 0
1998 1
1999 4
2000 8
2001 3
2002 9
2003 1
2004 1
2005 5
2006 5
2007 9
2008 4
2009 3
2010 9
2011 7
2012 9
2013 3
2014 1
2015 2
2016 3
2017 4
2018 1
2019 1
2020 3
2021 11
2022 9
2023 5
2024 9
2025 3
Total 135
Download as CSV

Ido Pen publication distribution in Ecology and Evolution in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Ecology and Evolution in 2026. The highlighted bar marks where Ido Pen sits on this spectrum.

No. of scientists
100 200 300
Bar chart with 100 bars. Horizontal axis: publications, 37–41 to 531+. Vertical axis: number of scientists, 0 to 333. Most scientists, 333, have 97–101 publications. The last bar groups every scientist with 531 publications or more. The highlighted bar, 107–111 publications, is where this scientist sits. 37–41 publications: 2 scientists 42–46 publications: 9 scientists 47–51 publications: 10 scientists 52–56 publications: 31 scientists 57–61 publications: 56 scientists 62–66 publications: 91 scientists 67–71 publications: 92 scientists 72–76 publications: 127 scientists 77–81 publications: 173 scientists 82–86 publications: 236 scientists 87–91 publications: 227 scientists 92–96 publications: 240 scientists 97–101 publications: 333 scientists 102–106 publications: 280 scientists 107–111 publications: 300 scientists 112–116 publications: 285 scientists 117–121 publications: 305 scientists 122–126 publications: 287 scientists 127–131 publications: 278 scientists 132–136 publications: 289 scientists 137–141 publications: 273 scientists 142–146 publications: 262 scientists 147–151 publications: 246 scientists 152–156 publications: 235 scientists 157–161 publications: 205 scientists 162–166 publications: 199 scientists 167–171 publications: 174 scientists 172–176 publications: 164 scientists 177–181 publications: 173 scientists 182–186 publications: 189 scientists 187–191 publications: 170 scientists 192–196 publications: 139 scientists 197–201 publications: 128 scientists 202–206 publications: 117 scientists 207–211 publications: 115 scientists 212–216 publications: 107 scientists 217–221 publications: 124 scientists 222–226 publications: 81 scientists 227–231 publications: 82 scientists 232–236 publications: 85 scientists 237–241 publications: 75 scientists 242–246 publications: 66 scientists 247–251 publications: 81 scientists 252–256 publications: 69 scientists 257–261 publications: 70 scientists 262–266 publications: 55 scientists 267–271 publications: 64 scientists 272–276 publications: 49 scientists 277–281 publications: 48 scientists 282–286 publications: 44 scientists 287–291 publications: 45 scientists 292–296 publications: 36 scientists 297–301 publications: 39 scientists 302–306 publications: 38 scientists 307–311 publications: 30 scientists 312–316 publications: 34 scientists 317–321 publications: 25 scientists 322–326 publications: 22 scientists 327–331 publications: 28 scientists 332–336 publications: 30 scientists 337–341 publications: 22 scientists 342–346 publications: 30 scientists 347–351 publications: 26 scientists 352–356 publications: 22 scientists 357–361 publications: 29 scientists 362–366 publications: 21 scientists 367–371 publications: 12 scientists 372–376 publications: 18 scientists 377–381 publications: 17 scientists 382–386 publications: 13 scientists 387–391 publications: 21 scientists 392–396 publications: 13 scientists 397–401 publications: 10 scientists 402–406 publications: 15 scientists 407–411 publications: 8 scientists 412–416 publications: 9 scientists 417–421 publications: 8 scientists 422–426 publications: 4 scientists 427–431 publications: 10 scientists 432–436 publications: 10 scientists 437–441 publications: 5 scientists 442–446 publications: 4 scientists 447–451 publications: 10 scientists 452–456 publications: 2 scientists 457–461 publications: 4 scientists 462–466 publications: 7 scientists 467–471 publications: 7 scientists 472–476 publications: 6 scientists 477–481 publications: 6 scientists 482–486 publications: 3 scientists 487–491 publications: 3 scientists 492–496 publications: 4 scientists 497–501 publications: 4 scientists 502–506 publications: 8 scientists 507–511 publications: 5 scientists 512–516 publications: 4 scientists 517–521 publications: 5 scientists 522–526 publications: 4 scientists 527–530 publications: 5 scientists 531+ publications: 100 scientists
37–41 publications 531+

This scientist: 108 publications — 24th percentile

24% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 531 publications or more.

View publications distribution as a table
Number of Ecology and Evolution scientists by publication count, Research.com 2026 ranking edition. Based on 8,381 ranked scientists.
Publications Scientists This scientist
37–41 2
42–46 9
47–51 10
52–56 31
57–61 56
62–66 91
67–71 92
72–76 127
77–81 173
82–86 236
87–91 227
92–96 240
97–101 333
102–106 280
107–111 300 108
112–116 285
117–121 305
122–126 287
127–131 278
132–136 289
137–141 273
142–146 262
147–151 246
152–156 235
157–161 205
162–166 199
167–171 174
172–176 164
177–181 173
182–186 189
187–191 170
192–196 139
197–201 128
202–206 117
207–211 115
212–216 107
217–221 124
222–226 81
227–231 82
232–236 85
237–241 75
242–246 66
247–251 81
252–256 69
257–261 70
262–266 55
267–271 64
272–276 49
277–281 48
282–286 44
287–291 45
292–296 36
297–301 39
302–306 38
307–311 30
312–316 34
317–321 25
322–326 22
327–331 28
332–336 30
337–341 22
342–346 30
347–351 26
352–356 22
357–361 29
362–366 21
367–371 12
372–376 18
377–381 17
382–386 13
387–391 21
392–396 13
397–401 10
402–406 15
407–411 8
412–416 9
417–421 8
422–426 4
427–431 10
432–436 10
437–441 5
442–446 4
447–451 10
452–456 2
457–461 4
462–466 7
467–471 7
472–476 6
477–481 6
482–486 3
487–491 3
492–496 4
497–501 4
502–506 8
507–511 5
512–516 4
517–521 5
522–526 4
527–530 5
531+ 100
Download as CSV

Ido Pen D-index placement in Ecology and Evolution in 2026

The chart shows the D-index (discipline H-index) distribution of Ecology and Evolution scientists ranked by Research.com in 2026. The highlighted bar marks where Ido Pen sits on this spectrum.

No. of scientists
50 100 150 200 250
Bar chart with 92 bars. Horizontal axis: D-Index, 30 to 121+. Vertical axis: number of scientists, 0 to 280. Most scientists, 280, have 40 D-Index. The last bar groups every scientist with 121 D-Index or more. The highlighted bar, 43 D-Index, is where this scientist sits. 30 D-Index: 95 scientists 31 D-Index: 123 scientists 32 D-Index: 191 scientists 33 D-Index: 245 scientists 34 D-Index: 252 scientists 35 D-Index: 244 scientists 36 D-Index: 249 scientists 37 D-Index: 244 scientists 38 D-Index: 254 scientists 39 D-Index: 254 scientists 40 D-Index: 280 scientists 41 D-Index: 274 scientists 42 D-Index: 266 scientists 43 D-Index: 233 scientists 44 D-Index: 248 scientists 45 D-Index: 208 scientists 46 D-Index: 201 scientists 47 D-Index: 213 scientists 48 D-Index: 207 scientists 49 D-Index: 172 scientists 50 D-Index: 209 scientists 51 D-Index: 199 scientists 52 D-Index: 153 scientists 53 D-Index: 169 scientists 54 D-Index: 163 scientists 55 D-Index: 148 scientists 56 D-Index: 120 scientists 57 D-Index: 146 scientists 58 D-Index: 137 scientists 59 D-Index: 140 scientists 60 D-Index: 104 scientists 61 D-Index: 119 scientists 62 D-Index: 118 scientists 63 D-Index: 99 scientists 64 D-Index: 77 scientists 65 D-Index: 100 scientists 66 D-Index: 86 scientists 67 D-Index: 83 scientists 68 D-Index: 75 scientists 69 D-Index: 94 scientists 70 D-Index: 57 scientists 71 D-Index: 68 scientists 72 D-Index: 59 scientists 73 D-Index: 63 scientists 74 D-Index: 66 scientists 75 D-Index: 60 scientists 76 D-Index: 42 scientists 77 D-Index: 49 scientists 78 D-Index: 39 scientists 79 D-Index: 37 scientists 80 D-Index: 43 scientists 81 D-Index: 41 scientists 82 D-Index: 45 scientists 83 D-Index: 34 scientists 84 D-Index: 40 scientists 85 D-Index: 35 scientists 86 D-Index: 55 scientists 87 D-Index: 24 scientists 88 D-Index: 24 scientists 89 D-Index: 30 scientists 90 D-Index: 23 scientists 91 D-Index: 25 scientists 92 D-Index: 27 scientists 93 D-Index: 27 scientists 94 D-Index: 13 scientists 95 D-Index: 23 scientists 96 D-Index: 12 scientists 97 D-Index: 14 scientists 98 D-Index: 19 scientists 99 D-Index: 16 scientists 100 D-Index: 14 scientists 101 D-Index: 5 scientists 102 D-Index: 21 scientists 103 D-Index: 16 scientists 104 D-Index: 10 scientists 105 D-Index: 7 scientists 106 D-Index: 7 scientists 107 D-Index: 7 scientists 108 D-Index: 6 scientists 109 D-Index: 9 scientists 110 D-Index: 11 scientists 111 D-Index: 12 scientists 112 D-Index: 9 scientists 113 D-Index: 7 scientists 114 D-Index: 3 scientists 115 D-Index: 8 scientists 116 D-Index: 5 scientists 117 D-Index: 11 scientists 118 D-Index: 4 scientists 119 D-Index: 2 scientists 120 D-Index: 5 scientists 121+ D-Index: 100 scientists
30 D-Index 121+

This scientist: 43 D-Index — 38th percentile

38% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 121 D-Index or more.

View D-Index distribution as a table
Number of Ecology and Evolution scientists by D-index, Research.com 2026 ranking edition. Based on 8,381 ranked scientists.
D-Index Scientists This scientist
30 95
31 123
32 191
33 245
34 252
35 244
36 249
37 244
38 254
39 254
40 280
41 274
42 266
43 233 43
44 248
45 208
46 201
47 213
48 207
49 172
50 209
51 199
52 153
53 169
54 163
55 148
56 120
57 146
58 137
59 140
60 104
61 119
62 118
63 99
64 77
65 100
66 86
67 83
68 75
69 94
70 57
71 68
72 59
73 63
74 66
75 60
76 42
77 49
78 39
79 37
80 43
81 41
82 45
83 34
84 40
85 35
86 55
87 24
88 24
89 30
90 23
91 25
92 27
93 27
94 13
95 23
96 12
97 14
98 19
99 16
100 14
101 5
102 21
103 16
104 10
105 7
106 7
107 7
108 6
109 9
110 11
111 12
112 9
113 7
114 3
115 8
116 5
117 11
118 4
119 2
120 5
121+ 100
Download as CSV

Overview

Ido Pen is affiliated with the University of Groningen in the Netherlands. Their research spans multiple fields primarily within Agricultural and Biological Sciences as well as Biochemistry, Genetics and Molecular Biology. The work focuses significantly on Ecology, Evolution, Behavior and Systematics, as well as Genetics, with considerable attention to specialized subfields including Ecology and Insect Science.

The scientist's research topics encompass a variety of areas related to animal and insect studies, including:

  • Animal Behavior and Reproduction
  • Plant and animal studies
  • Insect and Arachnid Ecology and Behavior
  • Evolutionary Game Theory and Cooperation
  • Wildlife Ecology and Conservation
  • Neurobiology and Insect Physiology Research
  • Genetic and Clinical Aspects of Sex Determination and Chromosomal Abnormalities

Ido Pen has contributed to multiple papers, some of which include:

  • Seasonal morphotypes of Drosophila suzukii differ in key life-history traits during and after a prolonged period of cold exposure, 2020, Ecology and Evolution
  • Asymmetry, division of labour and the evolution of ageing in multicellular organisms, 2021, Philosophical Transactions of the Royal Society B Biological Sciences
  • DNA methylation markers of age(ing) in non-model animals, 2023, Molecular Ecology
  • The evolution of social philopatry in female primates, 2020, American Journal of Physical Anthropology
  • Antagonistic pleiotropy and the evolution of extraordinary lifespans in eusocial organisms, 2021, Evolution Letters

Frequent coauthors collaborating with Ido Pen include:

  • Jan J. Kreider
  • Boris H. Kramer
  • Jan Komdeur
  • Ella Rees-Baylis
  • Leo W. Beukeboom

Their publications are often featured in journals such as:

  • bioRxiv (Cold Spring Harbor Laboratory)
  • Proceedings of the National Academy of Sciences
  • Evolution Letters
  • Journal of Animal Ecology
  • Journal of Evolutionary Biology

Best Publications

  • Cooperation and competition between relatives.

    Stuart A. West;Ido Pen;Ashleigh S. Griffin

  • Inclusive fitness theory and eusociality

    Patrick Abbot;Jun Abe;John Alcock;Samuel Alizon

  • Geographical patterns of adaptation within a species’ range: interactions between drift and gene flow

    M. Alleaume-Benharira;I. R. Pen;Ophélie Ronce

  • A Guide to Sexual Selection Theory

    Bram Kuijper;Ido Pen;Franz J. Weissing

  • An Evolutionary Perspective on Self-Organized Division of Labor in Social Insects

    Ana Duarte;Franz J. Weissing;Ido Pen;Laurent Keller

  • Sanctions and mutualism stability: when should less beneficial mutualists be tolerated?

    S. A. West;E. Toby Kiers;I. Pen;R. F. Denison

  • Adaptive sex allocation in birds: the complexities of linking theory and practice.

    Jan Komdeur;Ido Pen

  • Climate-driven population divergence in sex-determining systems

    Ido Pen;Tobias Uller;Barbara Feldmeyer;Anna Harts

  • Three epigenetic information channels and their different roles in evolution

    Nicholas Shea;Ido Pen;Tobias Uller

  • Towards a unified theory of cooperative breeding: the role of ecology and life history re-examined.

    Ido Pen;Franz J. Weissing

  • The evolution of sex ratios and sex-determining systems

    Tobias Uller;Ido Pen;Erik Wapstra;Leo W. Beukeboom

  • When is incomplete epigenetic resetting in germ cells favoured by natural selection

    Tobias Uller;Tobias Uller;Sinead English;Ido Pen

  • Genomic conflict in scale insects: the causes and consequences of bizarre genetic systems

    Laura Ross;Laura Ross;Ido Pen;David M. Shuker;David M. Shuker

  • Segregation distortion and the evolution of sex-determining mechanisms

    M.A. Kozielska;F. J. Weissing;L.W. Beukeboom;I.R. Pen

  • Optimal sex allocation: Steps towards a mechanistic theory

    Ido Pen;Franz J. Weissing

  • Caribbean mangroves and seagrass beds as daytime feeding habitats for juvenile French grunts, Haemulon flavolineatum

    Marieke C. Verweij;Ivan Nagelkerken;Suzanne L. J. Wartenbergh;Ido R. Pen

  • Sex ratios under asymmetrical local mate competition : Theory and a test with parasitoid wasps

    David M. Shuker;Ido Pen;Alison B. Duncan;Sarah E. Reece

  • The information value of non-genetic inheritance in plants and animals.

    Sinead English;Ido Pen;Nicholas Shea;Tobias Uller

  • Sex-ratio optimization with helpers at the nest

    Ido Pen;Franz J. Weissing

  • Sex ratio selection and multi‐factorial sex determination in the housefly: a dynamic model

    M.A. Kozielska;I.R. Pen;L.W. Beukeboom;F.J. Weissing

  • Parental energy expenditure in relation to manipulated brood size in the European kestrel Falco tinnunculus

    Charlotte Deerenberg;Ido Pen;Cornelis Dijkstra;Bart-Jan Arkies

Frequent Co-Authors

Franz J. Weissing
Franz J. Weissing University of Groningen
David M. Shuker
David M. Shuker University of St Andrews
Leo W. Beukeboom
Leo W. Beukeboom University of Groningen
Tobias Uller
Tobias Uller Lund University
Jan Komdeur
Jan Komdeur University of Groningen
Stuart A. West
Stuart A. West University of Oxford
Daniel J. C. Kronauer
Daniel J. C. Kronauer Rockefeller University
Erik Wapstra
Erik Wapstra University of Tasmania
Sarah E. Reece
Sarah E. Reece University of Edinburgh
Michael C. Singer
Michael C. Singer Plymouth University

External Links

Personal website of Ido Pen

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Best Scientists Citing Ido Pen

Stuart A. West

Stuart A. West

University of Oxford

Publications: 74

Andy Gardner

Andy Gardner

University of St Andrews

Publications: 63

Hanna Kokko

Hanna Kokko

Johannes Gutenberg University of Mainz

Publications: 42

Michael Taborsky

Michael Taborsky

University of Bern

Publications: 34

Tobias Uller

Tobias Uller

Lund University

Publications: 33

David M. Shuker

David M. Shuker

University of St Andrews

Publications: 33

Erik Wapstra

Erik Wapstra

University of Tasmania

Publications: 32

Rufus A. Johnstone

Rufus A. Johnstone

University of Cambridge

Publications: 28

Ashleigh S. Griffin

Ashleigh S. Griffin

University of Oxford

Publications: 25

Ivan Nagelkerken

Ivan Nagelkerken

University of Adelaide

Publications: 23

Franz J. Weissing

Franz J. Weissing

University of Groningen

Publications: 22

Jan Komdeur

Jan Komdeur

University of Groningen

Publications: 22

Laurent Lehmann

Laurent Lehmann

University of Lausanne

Publications: 22

Tamás Székely

Tamás Székely

University of Bath

Publications: 21

Martin A. Nowak

Martin A. Nowak

Harvard University

Publications: 21

E. Toby Kiers

E. Toby Kiers

Vrije Universiteit Amsterdam

Publications: 19

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