World's Best Scientists 2026 revealed!

D-Index & Metrics

Biology and Biochemistry

D-Index
101
Citations
41850
World Ranking
1426
National Ranking
813

Overview

Clarence A. Ryan was affiliated with Washington State University in the United States. Their research contributions spanned across Environmental Science and Engineering, with a particular focus on topics related to forest and ecosystem management.

The main fields of study associated with Ryan's work included:

  • Environmental Science
  • Engineering

Within these disciplines, Ryan explored multiple subfields, such as:

  • Global and Planetary Change
  • Mechanics of Materials
  • Atmospheric Science
  • Nature and Landscape Conservation
  • Mechanical Engineering

Ryan's research mostly concentrated on forest-related topics, specifically:

  • Forest Management and Policy
  • Fire Effects on Ecosystems
  • Forest Biomass Utilization and Management
  • Tree-ring Climate Responses
  • Forest Ecology and Management
  • Tree Root and Stability Studies

This focus was reflected in their publication record, which included papers in venues such as Forest Ecology and Management and Tree-Ring Research. Two notable articles published in 2025 were:

  • Redefining sustainable forestry for a climate emergency, 2025, Forest Ecology and Management
  • Looking at the Wood, Seeing the Trees and More: Australia-New Zealand Tree-Ring Science Conference, January 2025, Tree-Ring Research

Ryan collaborated frequently with several researchers, showing interdisciplinary engagement. Frequent co-authors included:

  • R. Alec Giffen
  • Frank Lowenstein
  • Robert T. Perschel
  • Mary L. Tyrrell
  • Gretel Boswijk

Best Publications

  • Interplant communication: airborne methyl jasmonate induces synthesis of proteinase inhibitors in plant leaves.

    Edward E. Farmer;Clarence A. Ryan

  • Wound-Induced Proteinase Inhibitor in Plant Leaves: A Possible Defense Mechanism against Insects.

    T. R. Green;C. A. Ryan

  • Octadecanoid Precursors of Jasmonic Acid Activate the Synthesis of Wound-Inducible Proteinase Inhibitors.

    Edward E. Farmer;Clarence A. Ryan

  • A Polypeptide from Tomato Leaves Induces Wound-Inducible Proteinase Inhibitor Proteins

    Gregory Pearce;Daniel Strydom;Scott Johnson;Clarence A. Ryan

  • Hydrogen Peroxide Acts as a Second Messenger for the Induction of Defense Genes in Tomato Plants in Response to Wounding, Systemin, and Methyl Jasmonate

    Martha L. Orozco-Cárdenas;Javier Narváez-Vásquez;Clarence A. Ryan

  • Hydrogen peroxide is generated systemically in plant leaves by wounding and systemin via the octadecanoid pathway

    Martha Orozco-Cardenas;Clarence A. Ryan

  • Expression of proteinase inhibitors I and II in transgenic tobacco plants: effects on natural defense against Manduca sexta larvae.

    Russell Johnson;Javier Narvaez;Gynheung An;Clarence Ryan

  • An endogenous peptide signal in Arabidopsis activates components of the innate immune response.

    Alisa Huffaker;Gregory Pearce;Clarence A. Ryan

  • SALICYLIC ACID INHIBITS SYNTHESIS OF PROTEINASE INHIBITORS IN TOMATO LEAVES INDUCED BY SYSTEMIN AND JASMONIC ACID

    S. H. Doares;J. Narvaez-Vasquez;A. Conconi;C. A. Ryan

  • Oligogalacturonides and chitosan activate plant defensive genes through the octadecanoid pathway.

    Steven H. Doares;Tatjana Syrovets;Elmar W. Weiler;Clarence A. Ryan

  • Regulation of Expression of Proteinase Inhibitor Genes by Methyl Jasmonate and Jasmonic Acid

    Edward E. Farmer;Russell R. Johnson;Clarence A. Ryan

  • An octadecanoid pathway mutant (JL5) of tomato is compromised in signaling for defense against insect attack.

    Gregg A. Howe;Jonathan Lightner;John Browse;Clarence A. Ryan

  • Structure, expression, and antisense inhibition of the systemin precursor gene

    Barry McGurl;Gregory Pearce;Martha Orozco-Cardenas;Clarence A. Ryan

  • Proteolytic Enzymes and Their Inhibitors in Plants

    C. A. Ryan

  • The cell surface leucine-rich repeat receptor for AtPep1, an endogenous peptide elicitor in Arabidopsis, is functional in transgenic tobacco cells.

    Yube Yamaguchi;Gregory Pearce;Clarence A. Ryan

  • Oligosaccharide Signals in Plants: A Current Assessment

    Clarence A. Ryan;Edward E. Farmer

  • Systemin activates synthesis of wound-inducible tomato leaf polyphenol oxidase via the octadecanoid defense signaling pathway.

    C P Constabel;D R Bergey;C A Ryan

  • PEPR2 Is a Second Receptor for the Pep1 and Pep2 Peptides and Contributes to Defense Responses in Arabidopsis

    Yube Yamaguchi;Alisa Huffaker;Anthony C. Bryan;Frans E. Tax

  • RALF, a 5-kDa ubiquitous polypeptide in plants, arrests root growth and development.

    Gregory Pearce;Daniel S. Moura;Johannes Stratmann;Clarence A. Ryan

  • Polypeptide signaling for plant defensive genes exhibits analogies to defense signaling in animals.

    Daniel R. Bergey;Gregg A. Howe;Clarence A. Ryan

Frequent Co-Authors

Gregory Pearce
Gregory Pearce Washington State University
Edward E. Farmer
Edward E. Farmer University of Lausanne
M. K. Walker-Simmons
M. K. Walker-Simmons United States Department of Agriculture
Gynheung An
Gynheung An Kyung Hee University
Vincent R. Franceschi
Vincent R. Franceschi Washington State University
Michael N. G. James
Michael N. G. James University of Alberta
Andreas Schaller
Andreas Schaller University of Hohenheim
Gregg A. Howe
Gregg A. Howe Michigan State University
John Browse
John Browse Washington State University
C. Peter Constabel
C. Peter Constabel University of Victoria

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