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Penny A. Handford

D-Index & Metrics

Biology and Biochemistry

D-Index
49
Citations
10020
World Ranking
17947
National Ranking
1416

Penny A. Handford publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Penny A. Handford sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 413 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 848 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 949 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 840 scientists 207–216 publications: 733 scientists 217–226 publications: 708 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 417 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 196 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 59 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 137 publications — 20th percentile

20% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Penny A. Handford D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Penny A. Handford sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 316 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 899 scientists 52–53 D-Index: 1,025 scientists 54–55 D-Index: 1,149 scientists 56–57 D-Index: 1,235 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,162 scientists 62–63 D-Index: 1,130 scientists 64–65 D-Index: 1,031 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 714 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 71 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 49 D-Index — 9th percentile

9% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Overview

Penny A. Handford is affiliated with the University of Oxford in the United Kingdom. Their research focuses on Biochemistry, Genetics, and Molecular Biology, with significant contributions in Molecular Biology, Cell Biology, Cancer Research, Immunology and Allergy, and Genetics.

Their work addresses several topics, including:

  • Developmental Biology and Gene Regulation
  • Protein Tyrosine Phosphatases
  • Retinal Development and Disorders
  • Cellular transport and secretion
  • RNA Research and Splicing
  • RNA modifications and cancer
  • Muscle Physiology and Disorders

Recent publications by Penny A. Handford include:

  • "A form of muscular dystrophy associated with pathogenic variants in JAG2," 2021, The American Journal of Human Genetics
  • "The Notch Signaling Pathway: Mechanistic Insights in Health and Disease," 2023, Engineering
  • "A form of muscular dystrophy associated with pathogenic variants in JAG2," 2021, The American Journal of Human Genetics
  • "An N-glycan on the C2 domain of JAGGED1 is important for Notch activation," 2022, Science Signaling
  • "The conserved C2 phospholipid-binding domain in Delta contributes to robust Notch signalling," 2021, EMBO Reports

They frequently publish in the following venues:

  • The American Journal of Human Genetics
  • Engineering
  • EMBO Reports
  • Science Signaling
  • PLoS ONE

Frequent co-authors collaborating with Penny A. Handford include:

  • Yao Meng
  • Bogusława Korona
  • Richard J. Suckling
  • Steven Johnson
  • Susan M. Lea

Best Publications

  • Somatic mutant clones colonize the human esophagus with age

    Iñigo Martincorena;Joanna C. Fowler;Agnieszka Wabik;Andrew R. J. Lawson

  • Solution structure of a pair of calcium-binding epidermal growth factor-like domains: implications for the Marfan syndrome and other genetic disorders.

    A.K Downing;V Knott;J.M Werner;C.M Cardy

  • The molecular genetics of Marfan syndrome and related disorders

    Peter N. Robinson;E. Arteaga-Solis;C. Baldock;G. Collod-Béroud

  • The structure of a Ca(2+)-binding epidermal growth factor-like domain: its role in protein-protein interactions.

    Zihe Rao;P. Handford;M. Mayhew;V. Knott

  • Key residues involved in calcium-binding motifs in EGF-like domains.

    Handford Pa;Mayhew M;Baron M;Winship Pr;Winship Pr

  • A conserved face of the Jagged/Serrate DSL domain is involved in Notch trans -activation and cis -inhibition

    Jemima Cordle;Steven Johnson;Joyce Zi Yan Tay;Pietro Roversi

  • The role of beta-hydroxyaspartate and adjacent carboxylate residues in the first EGF domain of human factor IX.

    D. J. G. Rees;I. M. Jones;P. A. Handford;S. J. Walter

  • Mutations in Fibrillin-1 Cause Congenital Scleroderma: Stiff Skin Syndrome

    B. L. Loeys;B. L. Loeys;E. E. Gerber;D. Riegert-Johnson;S. Iqbal

  • CRB1 mutation spectrum in inherited retinal dystrophies.

    Anneke I. den Hollander;Jason Davis;Saskia D. van der Velde-Visser;Marijke N. Zonneveld

  • Notch receptor-ligand binding and activation: insights from molecular studies.

    Chandramouli R. Chillakuri;Devon Sheppard;Susan M. Lea;Penny A. Handford

  • The first EGF-like domain from human factor IX contains a high-affinity calcium binding site.

    P A Handford;M Baron;M Mayhew;A Willis

  • Mutations in the TGFβ Binding-Protein-Like Domain 5 of FBN1 Are Responsible for Acromicric and Geleophysic Dysplasias

    Carine Le Goff;Clémentine Mahaut;Lauren W Wang;Slimane Allali

  • Microfibrils, elastin fibres and collagen fibres in the human intervertebral disc and bovine tail disc.

    Jing Yu;Uday Tirlapur;Jeremy Fairbank;Penny Handford

  • Solution structure of the transforming growth factor beta-binding protein-like module, a domain associated with matrix fibrils.

    X Yuan;A K Downing;V Knott;P A Handford

  • The CD46-Jagged1 interaction is critical for human TH1 immunity

    Gaëlle Le Friec;Devon Sheppard;Pat Whiteman;Christian M. Karsten

  • CALCIUM BINDING PROPERTIES OF AN EPIDERMAL GROWTH FACTOR-LIKE DOMAIN PAIR FROM HUMAN FIBRILLIN-1

    V Knott;A K Downing;C M Cardy;P Handford

  • Structural and functional properties of the human notch-1 ligand binding region.

    S Hambleton;N V Valeyev;A Muranyi;V Knott

  • Fibrillin: from domain structure to supramolecular assembly.

    P.A Handford;A.K Downing;D.P Reinhardt;L.Y Sakai

  • Allelic variation in normal human FBN1 expression in a family with Marfan syndrome: a potential modifier of phenotype?

    Sarah Hutchinson;Andre Furger;Dorothy Halliday;Daniel P. Judge

  • The Calcium Binding Properties and Molecular Organization of Epidermal Growth Factor-like Domains in Human Fibrillin-1

    P. Handford;A. K. Downing;Zihe Rao;D. R. Hewett

Frequent Co-Authors

Christina Redfield
Christina Redfield University of Oxford
Susan M. Lea
Susan M. Lea University of Oxford
George G. Brownlee
George G. Brownlee University of Oxford
Iain D. Campbell
Iain D. Campbell University of Oxford
Claudia Kemper
Claudia Kemper National Institutes of Health
Zihe Rao
Zihe Rao Tsinghua University
Simon N. Waddington
Simon N. Waddington University College London
Antony C. Willis
Antony C. Willis University of Oxford
Anne H. O’Donnell-Luria
Anne H. O’Donnell-Luria Broad Institute
Harry C. Dietz
Harry C. Dietz Johns Hopkins University School of Medicine

External Links

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Best Scientists Citing Penny A. Handford

Cay M. Kielty

Cay M. Kielty

Wellcome Centre for Cell-Matrix Research

Publications: 62

Dieter P. Reinhardt

Dieter P. Reinhardt

McGill University

Publications: 51

Claudia Kemper

Claudia Kemper

National Institutes of Health

Publications: 46

Lynn Y. Sakai

Lynn Y. Sakai

Oregon Health & Science University

Publications: 46

Bart Loeys

Bart Loeys

University of Antwerp

Publications: 41

Harry C. Dietz

Harry C. Dietz

Johns Hopkins University School of Medicine

Publications: 40

Peter N. Robinson

Peter N. Robinson

The Jackson Laboratory

Publications: 35

Johan Stenflo

Johan Stenflo

Lund University

Publications: 33

Robert S. Haltiwanger

Robert S. Haltiwanger

University of Georgia

Publications: 32

Jörg Hamann

Jörg Hamann

University of Amsterdam

Publications: 30

Douglas R. Keene

Douglas R. Keene

Shriners Hospitals for Children - Portland

Publications: 29

Robert P. Mecham

Robert P. Mecham

Washington University in St. Louis

Publications: 25

Daniel B. Rifkin

Daniel B. Rifkin

New York University

Publications: 25

Peter J. Campbell

Peter J. Campbell

Wellcome Sanger Institute

Publications: 23

Catherine Boileau

Catherine Boileau

Université Paris Cité

Publications: 23

Francesco Ramirez

Francesco Ramirez

Icahn School of Medicine at Mount Sinai

Publications: 23

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Andrea Gambotto

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