World's Best Scientists 2026 revealed!
João Ramalho-Santos

João Ramalho-Santos

D-Index & Metrics

Biology and Biochemistry

D-Index
56
Citations
12861
World Ranking
14311
National Ranking
57

João Ramalho-Santos publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where João Ramalho-Santos sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 196 publications — 48th percentile

48% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

João Ramalho-Santos D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where João Ramalho-Santos sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 56 D-Index — 28th percentile

28% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Overview

João Ramalho-Santos is affiliated with the University of Coimbra in Portugal and has contributed extensively to the fields of Medicine and Biochemistry, Genetics and Molecular Biology. Their research primarily addresses areas within Reproductive Medicine, Public Health, Environmental and Occupational Health, Molecular Biology, Pediatrics, Perinatology and Child Health, and Physiology.

The scientist's work focuses on topics such as Reproductive Biology and Fertility, Sperm and Testicular Function, Ovarian Function and Disorders, Birth, Development, and Health, Pluripotent Stem Cells Research, Reproductive System and Pregnancy, and CRISPR and Genetic Engineering.

Among their recent publications are the following papers:

  • Fluorescent probes for the detection of reactive oxygen species in human spermatozoa, 2020, Asian Journal of Andrology
  • Insights from qualitative research on NAFLD awareness with a cohort of T2DM patients: time to go public with insulin resistance?, 2020, BMC Public Health
  • The impact of antisperm antibodies on human male reproductive function: an update, 2021, Reproduction
  • Mitochondria-Targeted Compounds to Assess and Improve Human Sperm Function, 2021, Antioxidants and Redox Signaling
  • Isolation of Extracellular Vesicles from Human Follicular Fluid: Size-Exclusion Chromatography versus Ultracentrifugation, 2023, Biomolecules

Frequent co-authors collaborating with João Ramalho-Santos include:

  • Anabela Marisa Azul
  • Teresa Almeida-Santos
  • Ana Paula Sousa
  • Mireia Alemany i Pagès
  • Maria Soares

The scientist has published multiple times in prominent venues such as:

  • WIREs Mechanisms of Disease
  • European Journal of Clinical Investigation
  • Biomolecules
  • Reproduction
  • Methods in molecular biology

João Ramalho-Santos also has a record of book publications predominantly through Imprensa da Universidade de Coimbra eBooks. Notable titles include:

  • A Healthy Liver Will Always Deliver! (2020)
  • La prochaine fois, pensez à ton foie! (2020)
  • Um Fígado Equilibrado é Meio Caminho Andado! (2020)
  • ¡Un Hígado Equilibrado Es Medio Camino Andado! (2020)
  • Un Fegato Sano Ti Porta Lontano! (2020)
  • Eine gesunde Leber - nicht nur was für Streber (2020)
  • Zdrowa wątroba przy długim życiu Cię zachowa! (2020)
  • Sigues metge del teu propi fetge! (2020)
  • Zdravá játra vždy splní své povinnosti! (2020)
  • Folias Mitocondriais: Uma Breve Viagem Sobre a Energia da Vida (2025)

Best Publications

  • Ubiquitin tag for sperm mitochondria

    Peter Sutovsky;Ricardo D. Moreno;João Ramalho-Santos;Tanja Dominko

  • Energy metabolism in human pluripotent stem cells and their differentiated counterparts.

    Sandra Varum;Ana S. B. Rodrigues;Michelle B. Moura;Olga Momcilovic

  • Chd1 regulates open chromatin and pluripotency of embryonic stem cells

    Alexandre Gaspar-Maia;Adi Alajem;Fanny Polesso;Rupa Sridharan

  • Mitochondria functionality and sperm quality

    Alexandra Amaral;Bárbara Lourenço;Mónica Marques;João Ramalho-Santos

  • Ubiquitinated Sperm Mitochondria, Selective Proteolysis, and the Regulation of Mitochondrial Inheritance in Mammalian Embryos

    Peter Sutovsky;Ricardo D. Moreno;João Ramalho-Santos;Tanja Dominko

  • Mitochondrial functionality in reproduction: from gonads and gametes to embryos and embryonic stem cells

    João Ramalho-Santos;Sandra Varum;Sandra Amaral;Paula C Mota

  • Diabetes and the impairment of reproductive function: possible role of mitochondria and reactive oxygen species.

    Sandra Amaral;Paulo J Oliveira;João Ramalho-Santos

  • A putative, ubiquitin-dependent mechanism for the recognition and elimination of defective spermatozoa in the mammalian epididymis.

    P. Sutovsky;R. Moreno;J. Ramalho-Santos;T. Dominko

  • The combined human sperm proteome: cellular pathways and implications for basic and clinical science

    Alexandra Amaral;Judit Castillo;João Ramalho-Santos;Rafael Oliva

  • The Expression of Mitochondrial DNA Transcription Factors during Early Cardiomyocyte In Vitro Differentiation from Human Embryonic Stem Cells

    Justin C. St. John;João Ramalho-Santos;Heather L. Gray;Patti Petrosko

  • Human sperm tail proteome suggests new endogenous metabolic pathways

    Alexandra Amaral;Judit Castillo;Josep Maria Estanyol;José Luís Ballescà

  • Unique checkpoints during the first cell cycle of fertilization after intracytoplasmic sperm injection in rhesus monkeys.

    Laura Hewitson;Tanja Dominko;Diana Takahashi;Crista Martinovich

  • Effects of hyperglycemia on sperm and testicular cells of Goto-Kakizaki and streptozotocin-treated rat models for diabetes

    Sandra Amaral;António J. Moreno;Maria Sancha Santos;Raquel Seiça

  • Parabens in male infertility-is there a mitochondrial connection?

    Renata S. Tavares;Fátima C. Martins;Paulo J. Oliveira;João Ramalho-Santos

  • Ubiquitination of prohibitin in mammalian sperm mitochondria: possible roles in the regulation of mitochondrial inheritance and sperm quality control.

    Winston E. Thompson;João Ramalho-Santos;Peter Sutovsky

  • Enhancement of human embryonic stem cell pluripotency through inhibition of the mitochondrial respiratory chain.

    S. Varum;O. Momčilović;C. Castro;A. Ben-Yehudah

  • Identification of Proteins Involved in Human Sperm Motility Using High-Throughput Differential Proteomics

    Alexandra Amaral;Carla Paiva;Claudio Attardo Parrinello;Josep Maria Estanyol

  • The expression of polymerase gamma and mitochondrial transcription factor A and the regulation of mitochondrial DNA content in mature human sperm

    Alexandra Amaral;João Ramalho-Santos;Justin C. St John

  • Ca2+ Signals Generated by CatSper and Ca2+ Stores Regulate Different Behaviors in Human Sperm

    Wardah Alasmari;Sarah Costello;Joao Correia;Senga K. Oxenham

  • Not all sperm are equal: functional mitochondria characterize a subpopulation of human sperm with better fertilization potential.

    Ana Paula Sousa;Alexandra Amaral;Marta Baptista;Renata Tavares

Frequent Co-Authors

Gerald Schatten
Gerald Schatten University of Pittsburgh
Paulo J. Oliveira
Paulo J. Oliveira University of Coimbra
Peter Sutovsky
Peter Sutovsky University of Missouri
Nejat Düzgüneş
Nejat Düzgüneş University of the Pacific
Shlomo Nir
Shlomo Nir Hebrew University of Jerusalem
Denis Wirtz
Denis Wirtz Johns Hopkins University
António J.M. Moreno
António J.M. Moreno University of Coimbra
Gary M. Wessel
Gary M. Wessel Brown University
Rafael Oliva
Rafael Oliva University of Barcelona
Edward K. L. Chan
Edward K. L. Chan University of Florida

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