2026 Joseph E. Varner: Biology and Biochemistry Researcher – H-Index, Publications & Awards

Discipline name	D-Index	World Ranking	National Ranking	Publications	Citations
Biology and Biochemistry	62	10800	4690	142	13706

Discipline name

D-Index

World Ranking

National Ranking

Publications

Citations

Biology and Biochemistry

10800

4690

142

13706

Research.com Recognitions

1984 - Member of the National Academy of Sciences
1963 - Fellow of the American Association for the Advancement of Science (AAAS)

Overview

Joseph E. Varner was affiliated with Washington University in St. Louis in the United States. Their research work spanned multiple areas within medicine and immunology, with a particular focus on cancer and immune system interactions.

The scientist contributed significantly to the study of immune cells in cancer and cancer immunotherapy. Their main fields of study included Medicine and Immunology and Microbiology, with subfields covering Immunology, Oncology, Pulmonary and Respiratory Medicine, Molecular Biology, and Genetics.

Varner's research topics encompassed:

Immune cells in cancer
Cancer Immunotherapy and Biomarkers
Ferroptosis and cancer prognosis
Immune Cell Function and Interaction
Glioma Diagnosis and Treatment
PI3K/AKT/mTOR signaling in cancer
Cell Adhesion Molecules Research

The scientist's publications appeared frequently in notable journals and venues, with the highest number of papers in Cancer Research, followed by Cancer Immunology Research, Journal for ImmunoTherapy of Cancer, bioRxiv (Cold Spring Harbor Laboratory), and UNC Libraries.

The five recent papers published by Varner include:

"JAK inhibition enhances checkpoint blockade immunotherapy in patients with Hodgkin lymphoma" (2024, Science)
"PI3Kγ inhibition suppresses microglia/TAM accumulation in glioblastoma microenvironment to promote exceptional temozolomide response" (2021, Proceedings of the National Academy of Sciences)
"Eganelisib combined with immune checkpoint inhibitor therapy and chemotherapy in frontline metastatic triple-negative breast cancer triggers macrophage reprogramming, immune activation and extracellular matrix reorganization in the tumor microenvironment" (2024, Journal for ImmunoTherapy of Cancer)
"PI3Kγ maintains the self-renewal of acute myeloid leukemia stem cells by regulating the pentose phosphate pathway" (2024, Blood)
"PI3Kγ stimulates a high molecular weight form of myosin light chain kinase to promote myeloid cell adhesion and tumor inflammation" (2022, Nature Communications)

Over their career, frequent collaborators included Jaroslav Žák, Hui Chen, Marc Paradise, Anghesom Ghebremedhin, and John R. Teijaro, indicating active collaboration within the field.

Joseph E. Varner was recognized with membership in the National Academy of Sciences in 1984 and was a Fellow of the American Association for the Advancement of Science since 1963.

Best Publications

Gibberellic Acid-Enhanced Synthesis and Release of α-Amylase and Ribonuclease by Isolated Barley and Aleurone Layers

Maarten J. Chrispeels;J. E. Varner

864
Citations
Plant cell wall architecture

Joseph E. Varner;Joseph E. Varner;Liang-Shiou Lin;Liang-Shiou Lin

473
Citations
Gibberellic Acid-induced synthesis of protease by isolated aleurone layers of barley.

John V. Jacobsen;J. E. Varner

426
Citations
Hormonal Control of Enzyme Synthesis: On the Mode of Action of Gibberellic Acid and Abscisin in Aleurone Layers of Barley

Maarten J. Chrispeels;J. E. Varner

413
Citations
The bioassay of gibberellins

R L Jones;J E Varner

391
Citations
An Alternative Methylation Pathway in Lignin Biosynthesis in Zinnia

Zheng Hua Ye;Richard E. Kneusel;Ulrich Matern;Joseph E. Varner

387
Citations
Hydrogen peroxide and lignification

P.D. Olson;J.E. Varner

373
Citations
A test for de novo synthesis of enzymes: density labeling with H2O18 of barley alpha-amylase induced by gibberellic acid.

Philip Filner;J. E. Varner

363
Citations
Accumulation of hydroxyproline-rich glycoprotein mRNAs in response to fungal elicitor and infection

Allan M. Showalter;John N. Bell;Carole L. Cramer;John A. Bailey

358
Citations
An early response to gibberellic Acid not requiring protein synthesis.

Yosef Ben-Tal;J. E. Varner

333
Citations
HORMONAL CONTROL OF ENZYME SYNTHESIS IN BARLEY ENDOSPERM.

J. E. Varner;G. Ram Chandra

332
Citations
Immunocytolocalization of extensin in developing soybean seed coats by immunogold-silver staining and by tissue printing on nitrocellulose paper.

G I Cassab;J E Varner

329
Citations
Tissue-Specific Expression of Cell Wall Proteins in Developing Soybean Tissues.

Zheng-Hua Ye;Joseph E. Varner

315
Citations
An extracellular matrix protein in plants: characterization of a genomic clone for carrot extensin.

Jychian Chen;Joseph E. Varner

290
Citations
Environmental or developmental changes cause many enzyme activities of higher plants to rise or fall.

Philip Filner;Joseph E. Varner;John L. Wray

256
Citations
Isolation and characterization of cDNA clones for carrot extensin and a proline-rich 33-kDa protein

Jychian Chen;Joseph E. Varner

248
Citations
Reinforced Polyproline II Conformation in a Hydroxyproline-Rich Cell Wall Glycoprotein from Carrot Root

G J van Holst;J E Varner

231
Citations
Comparative localization of three classes of cell wall proteins.

Zheng-Hua Ye;Yan-Ru Song;Abraham Marcus;Joseph E. Varner

203
Citations
Induction of cysteine and serine proteases during xylogenesis in Zinnia elegans

Zheng-Hua Ye;Joseph E. Varner

203
Citations
Inhibition of Gibberellic Acid Induced Formation of α-Amylase by Abscisin II

Maarten J. Chrispeels;J. E. Varner

191
Citations

Frequent Co-Authors

Zheng-Hua Ye University of Georgia

Jacob Schaefer Washington University in St. Louis

Maarten J. Chrispeels University of California, San Diego

Ulrich Matern Philipp University of Marburg

Allan M. Showalter Ohio University

Barbara G. Pickard Washington University in St. Louis

Roger N. Beachy Washington University in St. Louis

Christopher J. Lamb John Innes Centre

John E. Heuser Washington University in St. Louis

If you think any of the details on this page are incorrect, let us know.

Joseph E. Varner

D-Index & Metrics

D-Index & Metrics

Biology and Biochemistry

Research.com Recognitions

Overview

Best Publications

Gibberellic Acid-Enhanced Synthesis and Release of α-Amylase and Ribonuclease by Isolated Barley and Aleurone Layers

Plant cell wall architecture

Gibberellic Acid-induced synthesis of protease by isolated aleurone layers of barley.

Hormonal Control of Enzyme Synthesis: On the Mode of Action of Gibberellic Acid and Abscisin in Aleurone Layers of Barley

The bioassay of gibberellins

An Alternative Methylation Pathway in Lignin Biosynthesis in Zinnia

Hydrogen peroxide and lignification

A test for de novo synthesis of enzymes: density labeling with H2O18 of barley alpha-amylase induced by gibberellic acid.

Accumulation of hydroxyproline-rich glycoprotein mRNAs in response to fungal elicitor and infection

An early response to gibberellic Acid not requiring protein synthesis.

HORMONAL CONTROL OF ENZYME SYNTHESIS IN BARLEY ENDOSPERM.

Immunocytolocalization of extensin in developing soybean seed coats by immunogold-silver staining and by tissue printing on nitrocellulose paper.

Tissue-Specific Expression of Cell Wall Proteins in Developing Soybean Tissues.

An extracellular matrix protein in plants: characterization of a genomic clone for carrot extensin.

Environmental or developmental changes cause many enzyme activities of higher plants to rise or fall.

Isolation and characterization of cDNA clones for carrot extensin and a proline-rich 33-kDa protein

Reinforced Polyproline II Conformation in a Hydroxyproline-Rich Cell Wall Glycoprotein from Carrot Root

Comparative localization of three classes of cell wall proteins.

Induction of cysteine and serine proteases during xylogenesis in Zinnia elegans

Inhibition of Gibberellic Acid Induced Formation of α-Amylase by Abscisin II

Frequent Co-Authors

Best Scientists Citing Joseph E. Varner

Related Biology and Biochemistry Rankings & Guides

2026 Is a Care Coordination Online Degree Worth It?

2026 Certified Nursing Assistant (CNA) vs. Registered Nurse (RN): Explaining the Difference

2026 State-Approved Online Healthcare Management Degree Master's Programs

2026 Are There Any One-Year Online Medical Billing & Coding Degree Programs Worth Considering?

2026 Is Psychiatric Mental Health Nurse Practitioner a Hard Major? What Students Should Know

2026 Cheapest and Shortest Online Ultrasound Technology Degree Programs That Pay Well: Cost, Duration, and Salary Outlook

2026 Psychiatric Nurse Practitioner vs. Therapist: Explaining the Difference

2026 What Does a PMHNP Do: Responsibilities, Requirements, and Salary

2026 Is Demand for Bioinformatics Degree Graduates Growing or Declining?

2026 Cheapest and Shortest Online Medical Assisting Degree Programs That Pay Well: Cost, Duration, and Salary Outlook

2026 How To Become a Marine Biologist? Salary & Career Paths

2026 Which Schools Offer Flexible Start-Anytime Enrollment for a Nursing Education Program?

2026 Military-Friendly Online Health Administration Degree Programs: Benefits, Accreditation, and Career Outcomes

2026 Easiest Online Healthcare Innovation Master's Degree Programs That Pay Well: High-Salary Degrees with Simple Admissions

2026 Health Administration Degree vs. Certificate: Which Should You Choose?

2026 Is Demand for Psychiatric Mental Health Nurse Practitioner Degree Graduates Growing or Declining?

2026 Fastest Online Biology Master's Degree Programs You Can Finish in Under 2 Years

2026 Family Nurse Practitioner vs. Psychiatric Mental Health Nurse Practitioner: Explaining the Difference

2026 Healthcare Management Degree vs. Certificate: Which Should You Choose?

2026 Medical Assistant vs. Medical Coder: Explaining the Difference

2026 Part-Time vs. Full-Time Ultrasound Technology Degree Programs

Recently Published Articles

2026 Most Affordable Online Physics Degree Programs

2026 SLP Application Checklist for First-Time Graduate School Applicants

2026 SLP Career Options in Home Health and Community Care

2026 Most Affordable Online Criminal Justice Degrees

2026 Best Online DNP Programs Without Clinicals

2026 How Many Credits Can You Transfer Into a Materials Science & Engineering Degree Program?

2026 SLP Career Growth From Clinical Practice to Leadership Roles

2026 The Best Undergraduate Majors for Future MSW Students

2026 How Many Credits Can You Transfer Into a Global Studies Degree Program?

2026 Applying to Online SLP Programs With a Low GPA: What to Know

2026 Fashion Design Master's Degree vs Doctorate: Career Paths & Salary Differences

2026 Best RN to BSN Programs With No Clinicals

2026 How Many Credits Can You Transfer Into an Interdisciplinary Studies Degree Program?

2026 Audio & Music Master's Degree vs Doctorate: Career Paths & Salary Differences

2026 Fastest Online Nutrition Degree Programs

2026 How Many Credits Can You Transfer Into a Health Education Degree Program?

2026 Substance Use Disorder Training in Online MSW Programs

2026 What to Expect From Your First Clinical Placement in SLP Graduate School

2026 Comparing Online SLP Programs by Faculty Expertise

2026 How Many Credits Can You Transfer Into a Social Media Degree Program?

2026 How Many Credits Can You Transfer Into a Healthcare Management Degree Program?