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Joseph E. Varner

D-Index & Metrics

Biology and Biochemistry

D-Index
62
Citations
13706
World Ranking
10800
National Ranking
4690

Joseph E. Varner publication distribution in Biology and Biochemistry in 2026

The chart shows the distribution of publications by all Research.com ranked scientists in the field of Biology and Biochemistry in 2026. The highlighted bar marks where Joseph E. Varner sits on this spectrum.

47–56 publications: 8 scientists 57–66 publications: 35 scientists 67–76 publications: 106 scientists 77–86 publications: 231 scientists 87–96 publications: 414 scientists 97–106 publications: 546 scientists 107–116 publications: 704 scientists 117–126 publications: 849 scientists 127–136 publications: 980 scientists 137–146 publications: 942 scientists 147–156 publications: 969 scientists 157–166 publications: 950 scientists 167–176 publications: 951 scientists 177–186 publications: 915 scientists 187–196 publications: 787 scientists 197–206 publications: 841 scientists 207–216 publications: 735 scientists 217–226 publications: 709 scientists 227–236 publications: 651 scientists 237–246 publications: 605 scientists 247–256 publications: 510 scientists 257–266 publications: 524 scientists 267–276 publications: 434 scientists 277–286 publications: 418 scientists 287–296 publications: 350 scientists 297–306 publications: 363 scientists 307–316 publications: 315 scientists 317–326 publications: 296 scientists 327–336 publications: 261 scientists 337–346 publications: 240 scientists 347–356 publications: 219 scientists 357–366 publications: 197 scientists 367–376 publications: 154 scientists 377–386 publications: 161 scientists 387–396 publications: 155 scientists 397–406 publications: 145 scientists 407–416 publications: 124 scientists 417–426 publications: 112 scientists 427–436 publications: 132 scientists 437–446 publications: 116 scientists 447–456 publications: 99 scientists 457–466 publications: 81 scientists 467–476 publications: 91 scientists 477–486 publications: 80 scientists 487–496 publications: 80 scientists 497–506 publications: 60 scientists 507–516 publications: 36 scientists 517–526 publications: 46 scientists 527–536 publications: 54 scientists 537–546 publications: 44 scientists 547–556 publications: 43 scientists 557–566 publications: 43 scientists 567–576 publications: 42 scientists 577–586 publications: 25 scientists 587–596 publications: 34 scientists 597–606 publications: 23 scientists 607–616 publications: 33 scientists 617–626 publications: 31 scientists 627–636 publications: 27 scientists 637–646 publications: 25 scientists 647–656 publications: 28 scientists 657–666 publications: 34 scientists 667–676 publications: 18 scientists 677–686 publications: 16 scientists 687–696 publications: 10 scientists 697–706 publications: 12 scientists 707–716 publications: 21 scientists 717–726 publications: 12 scientists 727–736 publications: 12 scientists 737–746 publications: 10 scientists 747–756 publications: 7 scientists 757–766 publications: 13 scientists 767–776 publications: 15 scientists 777–786 publications: 13 scientists 787–796 publications: 9 scientists 797–806 publications: 9 scientists 807–816 publications: 7 scientists 817–826 publications: 4 scientists 827–836 publications: 9 scientists 837–846 publications: 7 scientists 847–856 publications: 3 scientists 857–866 publications: 5 scientists 867–876 publications: 5 scientists 877–886 publications: 11 scientists 887–896 publications: 3 scientists 897–906 publications: 4 scientists 907–916 publications: 7 scientists 917–926 publications: 5 scientists 927–936 publications: 6 scientists 937–946 publications: 6 scientists 947–956 publications: 3 scientists 957–966 publications: 7 scientists 967–976 publications: 2 scientists 977–986 publications: 2 scientists 987–996 publications: 1 scientists 997–1,006 publications: 5 scientists 1,007–1,016 publications: 2 scientists 1,017–1,026 publications: 2 scientists 1,027 publications: 1 scientists 1,028+ publications: 100 scientists
47 publications 1,028+

This scientist: 142 publications — 23rd percentile

23% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 1,028 publications or more.

Joseph E. Varner D-index placement in Biology and Biochemistry in 2026

The chart shows the D-index (discipline H-index) distribution of Biology and Biochemistry scientists ranked by Research.com in 2026. The highlighted bar marks where Joseph E. Varner sits on this spectrum.

40–41 D-Index: 80 scientists 42–43 D-Index: 183 scientists 44–45 D-Index: 317 scientists 46–47 D-Index: 504 scientists 48–49 D-Index: 718 scientists 50–51 D-Index: 900 scientists 52–53 D-Index: 1,026 scientists 54–55 D-Index: 1,150 scientists 56–57 D-Index: 1,236 scientists 58–59 D-Index: 1,253 scientists 60–61 D-Index: 1,163 scientists 62–63 D-Index: 1,131 scientists 64–65 D-Index: 1,032 scientists 66–67 D-Index: 897 scientists 68–69 D-Index: 814 scientists 70–71 D-Index: 715 scientists 72–73 D-Index: 709 scientists 74–75 D-Index: 596 scientists 76–77 D-Index: 512 scientists 78–79 D-Index: 473 scientists 80–81 D-Index: 412 scientists 82–83 D-Index: 373 scientists 84–85 D-Index: 358 scientists 86–87 D-Index: 285 scientists 88–89 D-Index: 273 scientists 90–91 D-Index: 227 scientists 92–93 D-Index: 208 scientists 94–95 D-Index: 193 scientists 96–97 D-Index: 153 scientists 98–99 D-Index: 157 scientists 100–101 D-Index: 148 scientists 102–103 D-Index: 120 scientists 104–105 D-Index: 113 scientists 106–107 D-Index: 100 scientists 108–109 D-Index: 86 scientists 110–111 D-Index: 67 scientists 112–113 D-Index: 72 scientists 114–115 D-Index: 73 scientists 116–117 D-Index: 64 scientists 118–119 D-Index: 53 scientists 120–121 D-Index: 60 scientists 122–123 D-Index: 54 scientists 124–125 D-Index: 43 scientists 126–127 D-Index: 38 scientists 128–129 D-Index: 49 scientists 130–131 D-Index: 26 scientists 132–133 D-Index: 18 scientists 134–135 D-Index: 23 scientists 136–137 D-Index: 32 scientists 138–139 D-Index: 32 scientists 140–141 D-Index: 27 scientists 142–143 D-Index: 19 scientists 144–145 D-Index: 22 scientists 146–147 D-Index: 12 scientists 148–149 D-Index: 16 scientists 150–151 D-Index: 14 scientists 152–153 D-Index: 10 scientists 154–155 D-Index: 13 scientists 156–157 D-Index: 10 scientists 158–159 D-Index: 7 scientists 160–161 D-Index: 9 scientists 162–163 D-Index: 13 scientists 164–165 D-Index: 4 scientists 166 D-Index: 4 scientists 167+ D-Index: 98 scientists
40 D-Index 167+

This scientist: 62 D-Index — 46th percentile

46% of scientists in this discipline score the same or lower.

The last bar groups every scientist with 167 D-Index or more.

Research.com Recognitions

  • 1984 - Member of the National Academy of Sciences
  • 1963 - Fellow of the American Association for the Advancement of Science (AAAS)

Overview

Joseph E. Varner was affiliated with Washington University in St. Louis in the United States. Their research work spanned multiple areas within medicine and immunology, with a particular focus on cancer and immune system interactions.

The scientist contributed significantly to the study of immune cells in cancer and cancer immunotherapy. Their main fields of study included Medicine and Immunology and Microbiology, with subfields covering Immunology, Oncology, Pulmonary and Respiratory Medicine, Molecular Biology, and Genetics.

Varner's research topics encompassed:

  • Immune cells in cancer
  • Cancer Immunotherapy and Biomarkers
  • Ferroptosis and cancer prognosis
  • Immune Cell Function and Interaction
  • Glioma Diagnosis and Treatment
  • PI3K/AKT/mTOR signaling in cancer
  • Cell Adhesion Molecules Research

The scientist's publications appeared frequently in notable journals and venues, with the highest number of papers in Cancer Research, followed by Cancer Immunology Research, Journal for ImmunoTherapy of Cancer, bioRxiv (Cold Spring Harbor Laboratory), and UNC Libraries.

The five recent papers published by Varner include:

  • "JAK inhibition enhances checkpoint blockade immunotherapy in patients with Hodgkin lymphoma" (2024, Science)
  • "PI3Kγ inhibition suppresses microglia/TAM accumulation in glioblastoma microenvironment to promote exceptional temozolomide response" (2021, Proceedings of the National Academy of Sciences)
  • "Eganelisib combined with immune checkpoint inhibitor therapy and chemotherapy in frontline metastatic triple-negative breast cancer triggers macrophage reprogramming, immune activation and extracellular matrix reorganization in the tumor microenvironment" (2024, Journal for ImmunoTherapy of Cancer)
  • "PI3Kγ maintains the self-renewal of acute myeloid leukemia stem cells by regulating the pentose phosphate pathway" (2024, Blood)
  • "PI3Kγ stimulates a high molecular weight form of myosin light chain kinase to promote myeloid cell adhesion and tumor inflammation" (2022, Nature Communications)

Over their career, frequent collaborators included Jaroslav Žák, Hui Chen, Marc Paradise, Anghesom Ghebremedhin, and John R. Teijaro, indicating active collaboration within the field.

Joseph E. Varner was recognized with membership in the National Academy of Sciences in 1984 and was a Fellow of the American Association for the Advancement of Science since 1963.

Best Publications

  • Gibberellic Acid-Enhanced Synthesis and Release of α-Amylase and Ribonuclease by Isolated Barley and Aleurone Layers

    Maarten J. Chrispeels;J. E. Varner

  • Plant cell wall architecture

    Joseph E. Varner;Joseph E. Varner;Liang-Shiou Lin;Liang-Shiou Lin

  • Gibberellic Acid-induced synthesis of protease by isolated aleurone layers of barley.

    John V. Jacobsen;J. E. Varner

  • Hormonal Control of Enzyme Synthesis: On the Mode of Action of Gibberellic Acid and Abscisin in Aleurone Layers of Barley

    Maarten J. Chrispeels;J. E. Varner

  • The bioassay of gibberellins

    R L Jones;J E Varner

  • An Alternative Methylation Pathway in Lignin Biosynthesis in Zinnia

    Zheng Hua Ye;Richard E. Kneusel;Ulrich Matern;Joseph E. Varner

  • Hydrogen peroxide and lignification

    P.D. Olson;J.E. Varner

  • A test for de novo synthesis of enzymes: density labeling with H2O18 of barley alpha-amylase induced by gibberellic acid.

    Philip Filner;J. E. Varner

  • Accumulation of hydroxyproline-rich glycoprotein mRNAs in response to fungal elicitor and infection

    Allan M. Showalter;John N. Bell;Carole L. Cramer;John A. Bailey

  • An early response to gibberellic Acid not requiring protein synthesis.

    Yosef Ben-Tal;J. E. Varner

  • HORMONAL CONTROL OF ENZYME SYNTHESIS IN BARLEY ENDOSPERM.

    J. E. Varner;G. Ram Chandra

  • Immunocytolocalization of extensin in developing soybean seed coats by immunogold-silver staining and by tissue printing on nitrocellulose paper.

    G I Cassab;J E Varner

  • Tissue-Specific Expression of Cell Wall Proteins in Developing Soybean Tissues.

    Zheng-Hua Ye;Joseph E. Varner

  • An extracellular matrix protein in plants: characterization of a genomic clone for carrot extensin.

    Jychian Chen;Joseph E. Varner

  • Environmental or developmental changes cause many enzyme activities of higher plants to rise or fall.

    Philip Filner;Joseph E. Varner;John L. Wray

  • Isolation and characterization of cDNA clones for carrot extensin and a proline-rich 33-kDa protein

    Jychian Chen;Joseph E. Varner

  • Reinforced Polyproline II Conformation in a Hydroxyproline-Rich Cell Wall Glycoprotein from Carrot Root

    G J van Holst;J E Varner

  • Comparative localization of three classes of cell wall proteins.

    Zheng-Hua Ye;Yan-Ru Song;Abraham Marcus;Joseph E. Varner

  • Induction of cysteine and serine proteases during xylogenesis in Zinnia elegans

    Zheng-Hua Ye;Joseph E. Varner

  • Inhibition of Gibberellic Acid Induced Formation of α-Amylase by Abscisin II

    Maarten J. Chrispeels;J. E. Varner

Frequent Co-Authors

Zheng-Hua Ye
Zheng-Hua Ye University of Georgia
Jacob Schaefer
Jacob Schaefer Washington University in St. Louis
Maarten J. Chrispeels
Maarten J. Chrispeels University of California, San Diego
Ulrich Matern
Ulrich Matern Philipp University of Marburg
Allan M. Showalter
Allan M. Showalter Ohio University
Barbara G. Pickard
Barbara G. Pickard Washington University in St. Louis
Roger N. Beachy
Roger N. Beachy Washington University in St. Louis
Christopher J. Lamb
Christopher J. Lamb John Innes Centre
John E. Heuser
John E. Heuser Washington University in St. Louis

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Best Scientists Citing Joseph E. Varner

Russell L. Jones

Russell L. Jones

University of California, Berkeley

Publications: 50

Richard A. Dixon

Richard A. Dixon

University of North Texas

Publications: 43

Stephen C. Fry

Stephen C. Fry

University of Edinburgh

Publications: 30

Hiroo Fukuda

Hiroo Fukuda

University of Tokyo

Publications: 29

Pere Puigdomènech

Pere Puigdomènech

Spanish National Research Council

Publications: 25

John V. Jacobsen

John V. Jacobsen

Commonwealth Scientific and Industrial Research Organisation

Publications: 25

Christopher J. Lamb

Christopher J. Lamb

John Innes Centre

Publications: 19

Jacob Schaefer

Jacob Schaefer

Washington University in St. Louis

Publications: 18

Taku Demura

Taku Demura

Nara Institute of Science and Technology

Publications: 18

Derek T. A. Lamport

Derek T. A. Lamport

University of Sussex

Publications: 18

Maarten J. Chrispeels

Maarten J. Chrispeels

University of California, San Diego

Publications: 17

Marie-Thérèse Esquerré-Tugayé

Marie-Thérèse Esquerré-Tugayé

Paul Sabatier University

Publications: 17

Fang Chen

Fang Chen

University of North Texas

Publications: 16

Allan M. Showalter

Allan M. Showalter

Ohio University

Publications: 16

Geoffrey B. Fincher

Geoffrey B. Fincher

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Publications: 15

Keith Roberts

Keith Roberts

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Publications: 15

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